Systematics Of The Short-Tailed Whipscorpion Genus Stenochrus Chamberlin, 1922 (Schizomida: Hubbardiidae), With Descriptions Of Six New Genera And Five New Species Author Monjaraz-Ruedas, Rodrigo Author Prendini, Lorenzo Author Francke, Oscar F. text Bulletin of the American Museum of Natural History 2019 2019-06-20 2019 435 1 93 https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-2019/issue-435/0003-0090.435.1.1/Systematics-of-the-Short-Tailed-Whipscorpion-Genus-Stenochrus-Chamberlin-1922/10.1206/0003-0090.435.1.1.full journal article 10.1206/0003-0090.435.1.1 0003-0090 5416046 Key to Identification of the North American Genera of Hubbardiidae (Schizomida) 1. Cheliceral fixed finger with two teeth between two primary teeth, serrula composed of blunt teeth; pedipalp trochanter without prolateral spur; tarsal spurs symmetric; pygidial flagellum cylindrical, not dorsoventrally compressed ( ♂ ), with five flagellomeres ( ♀ ); spermathecae ( ♀ ) with one pair of lobes and without chitinized arch....... ................ Protoschizomidae Rowland, 1975 – Cheliceral fixed finger with three or more teeth between two primary teeth, serrula composed of several hyaline teeth (fig. 12); pedipalp trochanter with prolateral spur; tarsal spurs asymmetric; pygidial flagellum dorsoventrally compressed or globose ( ♂ ), with three or four flagellomeres ( ♀ ); spermathecae ( ♀ ) with two or more pairs of lobes and chitinized arch (figs. 8, 9).................................2 ( Hubbardiidae Cook, 1899 ) 2. Pedipalp femur, prolateral surface with four setae ( Fvr ); opisthosomal segment XII ( ♂ ) with conspicuous posterodorsal process; spermathecae ( ♀ ) with more than three pairs of extremely short, broad, and rounded lobes, clustered in groups.............. ................................... Hubbardia Cook, 1899 – Pedipalp femur, prolateral surface with three setae ( Fvr ); opisthosomal segment XII ( ♂ ) with or without inconspicuous posterodorsal process; spermathecae ( ♀ ) usually with two or four pairs of long, filiform lobes, not clustered in groups (figs. 8–10)...................3 3. Opisthosomal segment II with four setae; pedipalp femur ( ♂ ), setae Fv 1 and Fv 2 forming spiniform setiferous tubercles; prolateral tarsal spur ( ♂ ) present; pygidial flagellum ( ♀ ) with four flagellomeres................. Mayazomus Reddell and Cokendolpher, 1995 – Opisthosomal segment II with two setae; pedipalp femur ( ♂ ), setae Fv 1 and Fv 2 acuminate, setiform or spiniform; prolateral tarsal spur ( ♂ ) absent; pygidial flagellum ( ♀ ) with three flagellomeres....................4 4. Cheliceral movable finger with lamella or teeth (fig. 12B, C)...........................................5 – Cheliceral movable finger smooth (fig. 12A).......8 5. Opisthosomal segment XII with small, inconspicuous posterodorsal process ( ♂ ); pygidial flagellum ( ♂ ) usually globose; spermathecae ( ♀ ), lateral lobes greatly reduced (usually absent) compared to median lobes, and usually with apical bulbs............................................................ Pacal Reddell and Cokendolpher, 1995 – Opisthosomal segment XII without posterodorsal process ( ♂ ); pygidial flagellum ( ♂ ) dorsoventrally compressed; spermathecae ( ♀ ), lateral and median lobes subequal, without apical bulbs................................6 6. Opisthosomal segments VIII–XI ( ♂ ) elongated (fig. 18 C ) ; pygidial flagellum ( ♂ ) elongated with posterior constriction in dorsal view (fig. 19D–F); spermathecae ( ♀ ), chitinized arch mug shaped (fig. 8E, F).......... Heteroschizomus Rowland, 1973 , stat. rev. – Opisthosomal segments VIII–XI ( ♂ ) not elongated (fig. 15); pygidial flagellum ( ♂ ) elliptical or trilobed (figs. 19G–I, 23G–I); spermathecae ( ♀ ), chitinized arch wih lateral tips projected and acute (fig. 9A, B, F)..............7 7. Cheliceral movable finger with lamella (fig. 12C); pedipalp patella with five Pm setae, tibia setal formula 5-5-6 (fig. 14C); pygidial flagellum ( ♂ ) with pair of dorsosubmedian circular depressions, without projections (fig. 19G–I); spermathecae ( ♀ ), lateral and median lobes equal, with several apical sclerotizations in both pairs of lobes, creating appearence of leafy tree (fig. 9A, B) ............................................ Nahual , gen. nov. – Cheliceral movable finger serrate, comprising several small teeth (fig. 13I); pedipalp patella with four Pm setae, tibia setal formula 3-3-4 (fig. 14D); pygidial flagellum ( ♂ ) without depressions and with pair of dorsosubmedian projections (fig. 23G–I); spermathecae ( ♀ ), lateral lobes reduced to approximately 3/4 the length of median lobes, without apical sclerotizations in both pairs of lobes (fig. 9F)............................. ......................... Troglostenochrus , gen. nov. 8. Propeltidium with three pairs of dorsal setae (fig. 11A)......................................................9 – Propeltidium with two pairs of dorsal setae (fig. 11B)....................................................11 9. Pedipalp tibia with prominent lateral spur; spermathecae ( ♀ ) with four asymmetric pairs of hand-shaped lobes; pygidial flagellum ( ♂ ) globose, with medial depression.................................. Sotanostenochrus Reddell and Cokendolpher, 1991 – Pedipalp tibia without lateral spur; pygidial flagellum ( ♂ ) deltoid/spear shaped (figs. 16D–F, 19A– C ), with paired submedian depressions ; spermathecae ( ♀ ) with two pairs of filiform lobes..........................10 10. Pedipalp tibia, setal formula 3-3-5 (fig. 14B); pygidial flagellum ( ♂ ) bulbous/spear shaped, with paired submedian depressions not fused posteriorly (fig. 19A– C ) ; spermathecae ( ♀ ), median lobes parenthesis shaped, apex laterally directed, median and lateral lobe bases aligned, chitinized arch U-shaped (fig. 9 C )............................. ............................... Schizophyxia , gen. nov . – Pedipalp tibia, setal formula 4-3-5 (fig. 14A); pygidial flagellum ( ♂ ) deltoid (fig. 16A–F), with paired posterior depressions fused posteriorly; spermathecae ( ♀ ), median lobes straight, apex vertically directed, median lobe bases posterior to lateral lobe bases, chitinized arch inverse arch shaped (fig. 8A)............... Ambulantactus , gen. nov. 11. Pedipalps ( ♂ ) robust; pedipalp femur ( ♂ ), setae Fv 1 and Fv 2 forming spiniform setiferous tubercles, Fe 1 , Fe 5 , Ter 7 and Tir 5 spiniform; pygidial flagellum ( ♂ ) elliptical to lanceolate, with pair of dorsosubmedian depressions fused posteriorly; spermathecae ( ♀ ), lateral lobes swollen, drop shaped....... ............................ Olmecazomus , nom. nov. – Pedipalps ( ♂ ) homeomorphic; pedipalp femur ( ♂ ), setae Fe 1 , Fe 5 , Fv 1 , Fv 2 , Ter 7 and Tir 5 acuminate, setiform; pygidial flagellum ( ♂ ) ovate, subrhomboidal, or bulbous (figs. 16G–I, 23D–F), with pair of dorsosubmedian pits or depressions not fused posteriorly; spermathecae ( ♀ ), lateral lobes slender, filiform (figs. 8B, D)........................12 12. Pedipalp trochanter, apical process with bump (fig. 7C, H); pygidial flagellum ( ♂ ) subrhomboidal, and acuminate or lanceolate posteriorly (fig. 19A– C ), with pair of shallow dorsosubmedian pits ; spermathecae ( ♀ ), lateral lobes at least 2/3 the length of, or longer than, median lobes, and with apical bulbs (fig. 8 C , D).................................. ................................ Harveyus , gen. nov . – Pedipalp trochanter, apical process acute or absent (fig. 7B); pygidial flagellum ( ♂ ) elliptical, cordate or bulbose, and not acuminate or lanceolate posteriorly (fig. 23D–F), with pair of deep dorsosubmedian pits or without dorsal depressions; spermathecae ( ♀ ), lateral lobes at most half the length of median lobes, and without apical bulbs (figs. 8B, 9E).....................................13 13. Pygidial flagellum ( ♂ ) ovate or cordate, with pair of dorsosubmedian pits embedded in dorsomedian depression (fig. 20B); spermathecae ( ♀ ), median lobes similar width to lateral lobes (fig. 10 C ), curved laterally (J-shaped or parenthesis shaped) and sclerotized apically, median lobe bases posterior to lateral lobe bases (figs. 9E, 10 C )............ .................. Stenochrus Chamberlin, 1922 – Pygidial flagellum ( ♂ ) bulbous, without dorsal depressions or pits (fig. 16G–I); spermathecae ( ♀ ), median lobes broader than lateral lobes, slightly curved laterally and apically, and unsclerotized apically (fig. 8B), median and lateral lobe bases aligned (fig. 10A) ............................................ Baalrog , gen. nov.