Review of the genus Typhloiulus Latzel, 1884 in the Dinaric region, with a description of four new species and the first description of the male of Typhloiulus insularis Strasser, 1938 (Diplopoda: Julida: Julidae) Author Antić, Dragan Ž. Author Dražina, Tvrtko Author Rađa, Tonći Author Lučić, Luka R. Author Makarov, Slobodan E. text Zootaxa 2018 2018-08-02 4455 2 258 294 journal article 29106 10.11646/zootaxa.4455.2.2 431a0907-2b69-4c52-86c6-6a0eb0c04d2e 1175-5326 1457167 595A9780-C604-465C-BDE2-589B87A4DB52 Typhloiulus insularis Strasser, 1938 Figs 5 , 6 , 18C Material studied (total: 2 males, 1 female). Topotypes : 1 male , CROATIA , Island of Cres , Beli , Petričevići , Čampari Pit ( 45.123881°N , 14.333628°E ), 8 April 2001 , R. Ozimec & B. Jalžić leg. (IZB) ; 1 male , same data except: 7 April 2012 , T. Čuković leg. ( CBSS ) ; 1 female , same data except: 20 October , 2000, R. Ozimec & B. Jalžić leg. ( CSBB ) . Type locality. CROATIA: Island of Cres, Čampari Pit. Diagnosis. This species is most similar to some members of the subgenus Stygiiulus , such as T. illyricus , T. maximus Verhoeff, 1929 , and T. montellensis Verhoeff, 1930 (see Discussion). All these species are characterized by the presence of a stongly developed, fimbriated velum with an acuminate, posterior thickening. Typhloiulus insularis differs clearly from these three species in the presence of a much smaller body and in certain details of gonopod structure, such as the presence of a more robust opisthomere without a posterior node and the presence of a well-developed, spoon-shaped mesomere. Redescription. Based on topotypes (males) 10.5–12 mm long, vertical diameter of the largest body ring 0.55–0.65 mm , body with 36 or 43 podous rings + 1 or 2 apodous rings + telson. Topotype female 15.5 mm long, vertical diameter of the largest body ring 0.75 mm , body with 45 podous rings + 1 apodous ring + telson. Colouration : ( Figs 5A , 18C ). Depigmented, yellowish white. Head : ( Fig. 5B ). Labrum with three labral teeth, with 4 supralabral and 5+5 labral setae in smaller topotype male. Gnathochilarium with promentum kite-shaped, elongated; lamellae linguales with 2+2 setae in one row, stipites each with 3 setae. Antennae 1 mm long in smaller topotype male, its length 180% of vertical diameter of the largest body ring. Length of antennomeres I–VIII (in mm): 0.06 (I), 0.2 (II), 0.15 (III), 0.15 (IV), 0.2 (V), 0.13 (VI), 0.08 (VII), and 0.03 (VIII). Length/width ratio of antennomeres I–VII: 0.9 (I), 2.5 (II), 2 (III), 1.6 (IV), 1.8 (V), 1.3 (VI), and 1.3 (VII). Antennomeres V and VI each with a terminal corolla of large bacilliform sensilla; antennomere VII with a terminal corolla of microsetae. Body rings : ( Fig. 5C ). Barely visible metazonal longitudinal striations. Midbody rings with ca. 10 short metazonal setae. Length of midbody setae ca. 7% of vertical diameter of rings. Telson : ( Fig. 5E, F ). Epiproct with 2+2 dorsal and 6+6 ventro-lateral, long setae. Preanal process short, acuminate, with a small ventral protuberance and with two lateral setae. Paraprocts rounded, with numerous long setae. Hypoproct subrounded, with 1+1 long apical setae. Male legs : First pair of legs modified, hook-shaped ( Fig. 6A ); only three podomeres visible. Coxa without setae; prefemur each with two setae; femur, postfemur, and tibiotarsus coalesced, with two setae. Tips globular and tuberculate. Second pair of legs without ventral pads. Penis : ( Fig. 6B ). Bilobed. Lobes short, separated by a Vshaped sinus. Ventral margin of body ring 7 : ( Fig. 5D ). Poorly-developed, subtrapezoid. Gonopods : ( Fig. 6C–E ). Promere ( p ) and mesomere ( m ) shorter than opisthomere ( o ). Promere ( p ) spatulate, shorter than m , mesal lobe ( ml ) high and wide, without setae; telopodite ( t ) small, globular. Mesomere ( m ) robust, spoon-shaped, with anterior corolla of denticles. Apical part of m covering apex of p in situ . Opisthomere ( o ) robust, characterized by the presence of a stongly developed, fimbriated velum ( v ) with an acuminate, posterior thickening. Mesally with one elongated spine. Solenomere ( s ) rather wide. Distribution. Croatia : Island of Cres, Petričevići, Beli, Čampari Pit ( type locality) ( Strasser 1938; present study ). Up to now, this species was known only from its type locality ( Fig. 21 , red circle). This is the first record of the species after its original description, which was based on a female, and the first record of males. Notes. Typhloiulus insularis is known only from a pit.