A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae) Author Sabu, Thomas K. Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com sabukthomas@gmail.com Author Abhitha, Prabhakar Litter Entomology Research Unit, P. G. & Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources & Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com Author Zhao, Danyang text Zootaxa 2008 2008-04-09 1745 1 30 46 https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3 journal article 10.11646/zootaxa.1745.1.3 1175­5334 5108342 Key to the genera of the tribe Physocrotaphini 1. Ligula prominent and distinctly longer than paraglossae ( Fig. 2 ); post-orbital surface area (gena) densely setose, with more than 40 setae and without tubercles; maxillary palpomere 4 shorter than 3; TLA = 24.5– 27.5 ( Fig. 23 ) ............................................................................................................. Helluodes Westwood. - Ligula truncate and distinctly shorter than paraglossae ( Figs. 3–5 ); post-orbital surface area (gena) sparsely setose with less than 10 setae and with or without tubercles; maxillary palpomere 4 longer than 3; TLA = 19.5 ................................................................................................................................................ 2 2. Mentum with bifid median tooth; ligula stout and of uniform width from base to apex ( Fig. 3 ); male sternite 3 ( Fig. 15 ) with single oval depression bearing filamentous elongated setae; TLA = 19–19.5 ( Fig. 26 ) ................................................................................................................................... Physocrotaphus Parry. - Mentum with triangular median tooth; ligula slender and gradually widening towards apex ( Figs. 4, 5 ); male sternite 3 with two oval depressions bearing filamentous elongated setae ( Fig. 16 )..........................3 3. Antennae longer, extended beyond the base of pronotum by 2–3 antennomeres; dorsal and ventral surfaces of antennomeres 5–11 devoid of impressions at base; antennomeres 2–11 with apical ring of elongate setae; mandibles long, narrow and weakly arcuate; anterior transverse sulcus of pronotum weakly developed; TLA = 7–17.6 ( Fig. 24 ) .................................................................... Pogonoglossus Chaudoir. - Antennae shorter, extended beyond the base of pronotum by only the last; dorsal and ventral surfaces of antennomeres 5–11 bi-impressed at base; antennomeres 2–11 without apical ring of setae; mandibles short, wide and weakly arcuate; anterior transverse sulcus of pronotum very deep; length 14.5–16.8mm . ( Fig. 25 ) ............................................................................................................................. Schuelea Baehr.