A review of Australian long-horned caddisflies in the Oecetis pechana-group (Trichoptera: Leptoceridae), with descriptions of thirteen new species Author Wells, Alice text Memoirs of Museum Victoria 2006 2006-12-31 63 2 107 128 https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-63-issue-2-2006/pages-107-128/ journal article 10.24199/j.mmv.2006.63.13 1447-2554 8064969 Oecetis quadrula sp. nov. Figures 40–42, 70 Material examined. Holotype . Male, NT , Kakadu National Park , UDP Falls , 7 Sep 1979 , J. Blyth ( NMV T-18844, on slide). Paratypes . NT: 1 male , Litchfield National Park , Florence Falls , 9 Apr 1991 , Horak , Upton and Wells ( ANIC ) ; 1 male , Jabiru Town Lake , 13 May 1991 , C. Humphrey ( ANIC ) ; 1 male , Kakadu National Park , Baroalba Springs , 4 Oct 1991 , Wells and Webber ( NTM ) ; 1 male , same locality, 16 Jan 1992 , Wells , Webber and Bickel ( NTM ) ; 1 male , Magela Creek Site 009, 3 Mar 1992 , A. Wells ( NTM ) . Other material. NT: 1 male , Muirella Park , 12 Oct 1972 , J.C. Cardale ( ANIC ) ; 1 male , 12°17'S , 133°13'E , Birraduk Ck , 18 km E by N of Oenpelli , 4–5 Jun 1973 , J.C. Cardale ( ANIC ) ; 1 male , 12°31'S , 132°54'E , 9 km N by E of Mudginberri HS, 10–11 Jun 1973 , J.C. Cardale ( ANIC ) ; 1 male , 12°50'S , 132°51'E , 16 km E by N of Mt Cahill , 16 Jun 1973 , J.C. Cardale ( ANIC ) ; 1 male , 16°25'S , 136°05'E , 45 km SW by S of Borroloola , 5 Nov 1975 , J.C. Cardale ( ANIC ) ; male, Jim Jim Creek , 3 km below falls, Kakadu National Park , 1 Sep 1979 , J. Blyth ( NMV , on slide) ; male, South Alligator R. at UDP Falls [Gunlom], 7 Sep 1979 , J. Blyth ( NMV , on slide) ; 1 male , ARR, creek 5 km W of OSS South Alligator Field Station , 19 Apr 1989 , Wells and Suter ( NMV ) ; 1 male , Melville I. , Andranangoo Creek , 11°31'S , 130°54'E , 8 Oct 1996 , G.R. Brown ( NTM ) ; 1 male , Melville I. , Maxwell Creek , 11°33'S , 130°35'E , 14 Oct 1996 , G.R. Brown ( NTM ) ; 1 male , same locality and collector, 15 Oct 1996 ( NTM ) . WA: numerous males, females, 15°02'S , 126°55'E , Drysdale R., 3–5 Jul 1975 , I.B.F. Common ( ANIC ) ; numerous males, females, 15°02'S , 126°55'E , Drysdale R., 3–8 Aug 1975 , I.B.F. Common and M.S. Upton ( ANIC ) ; male, Mitchell Plateau , Camp Creek , 31 Jan 1978 , J.E. Bishop ( NMV , on slide) ; male, Mitchell Plateau , Camp Creek at Crusher , 15 Feb 1979 , J.E. Bishop ( NMV , on slide) ; 3 males , Mitchell Plateau , at Mining Camp , 14°49'S , 125°50'E , 9–19 May 1983 , J.C. Cardale ( ANIC ) ; 1 male , Kimberley , Mt Barnett Station , Manning Gorge , 30 Apr 1992 , P.S. Cranston ( ANIC ) ; 1 male , 15°55'S , 127°56'E , El Questro Station , Chamberlain R. pool, 26 Aug 1996 , I. Edwards , NMV ); numerous males, females, 15°42'S , 126°22'E , Drysdale R. Crossing , Kalumburu Road , 28 Aug 1996 , I. Edwards ( NMV ) ; 2 males , 14°54'S , 126°12'E , King Edward R., 3 Sep 1996 , I. Edwards ( NMV ) ; 1 male , 16°40'S , 125°56'E , Manning R. near Mt Barnett , 8 Sep 1996 , I. Edwards ( NMV ) . Qld : 1 male , The Crater , Atherton Tableland , 25 Apr 1970 , S.R. Curtis ( ANIC ) ; numerous males, females, Tewah Creek , Tin Can Bay , 17–18 Oct 1971 , S.R. Monteith ( ANIC ) ; 1 male , Upper Freshwater Creek , Whitfield Range near Cairns , 15 Dec 1974 , Moulds ( NMV , on slide) ; 1 male , Mulgrave R. W of Gordonvale , 29 Apr 1979 , Wells ( NMV ) ; 1 male , 15°12'S , 143°52'E , Hann R. NW by W of Laura , 27 Jun 1986 , J.C. Cardale ( ANIC ) . NSW: 4 males , 8 females , Wallagarough R. , Princes Highway bridge, 29 Jan 1975 , A. Neboiss ( NMV ) . Vic .: 1 male , Wingan R., 8 km S Princes Hwy , 30 Jan 1975 , A. Neboiss ( NMV , slide) ; 1 male , Tanjil R. near Old Tanjil, S 34, Latrobe C. Survey , 5 Feb 1980 ( NMV ) . Diagnosis. In the male, resembling O. australis and O. burtoni in having long downy vestiture on the wings and no scales, but distinguished by absence of acute ventro-mesial processes on the inferior appendages, and inferior appendages short and stout in ventral view, tapered distally in lateral view, phallus short and beak-like, with a short slightly curved paramere. The form of the inferior appendages of this species is similar to O. jenniae , but the phallus is far shorter, its paramere short and slender. Description . Male. Wings rounded apically, with long hair on veins; forewing without scales, length 4.6–7.0 mm. Antennae with a brush of long setae on leading edge of segments. Abdominal segments 3–5 with tergites sclerotised; segment IX narrow, lateral angles produced, segment X elongate coneshaped in ventral view, in lateral view elongate, rounded apically and with a small triangular ventral process. Genitalia, see figs 40–42. Pre-anal appendages short, rounded, phallus scarcely longer than wide, with membranous apex of aedeagus folded and down-turned; a short, slightly curved paramere present. Inferior appendages in ventral view subquadrate, with apico-lateral lobes short, convergent; in lateral view, simple, straight, tapered at apex, without basi-dorsal pouch. Distribution. Found across northern Australia , from WA, NT and Qld, and from south-eastern Vic. Remarks. With a dense brush of setae on the leading edge of the antennae, antennae erect (at least in spirit material), and long maxillary palpi, males of this species look particularly moth-like. Material here referred to O. quadrula shows variability in genitalic form, but a conservative approach is taken in assigning all to a single species as examination of more material from the south-east of the continent is required to justify recognition of two distinct morpho-species. Etymology : Derived from the Latin, quadra — square, for the shape of the inferior appendages in ventral view.