Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda Author Neubauer, Thomas A. Author Harzhauser, Mathias Author Georgopoulou, Elisavet Author Mandic, Oleg Author Kroh, Andreas text Zootaxa 2014 2014-04-04 3785 3 453 468 journal article 5666 10.11646/zootaxa.3785.3.7 8eddca4b-08a5-433a-8e3a-c89fb483818e 1175-5326 4913363 D6E3D055-4390-4DC2-A9AC-0DBC40C2740F Theodoxus ( Calvertia ) brusinai ( Pavlović, 1931 ) Neritodonta brusinai Pavlović 1931: 27 , pl. 11, figs 21–32 [non Neritina brusinai Andrusov, 1909 ]. Type locality. Valley of Voljavča brook near Bresnica, Moravica, Serbia . Age. Pontian (= upper Messinian-lower Zanclean). Holotype . Natural History Museum , Belgrade , coll. no. 3301 ( Milošević 1962 , p. 29). Discussion. The status as a (secondary) homonym depends on the controversial generic and subgeneric affiliations of the involved species. Pavlović (1931 , p. 27) introduced this species within the genus Neritodonta Brusina, 1884 (see also Milošević 1962 ). As stated above, Neritodonta is a junior synonym of Calvertia Bourguignat, 1880 , which is treated as a subgenus of Theodoxus (see also Wenz 1929, p. 2934). The older species was introduced as Neritina brusinai Andrusov, 1909 and newly combined as Theodoxus ( Ninnia ) brusinai by Wenz (1929, p. 2986). Therefore, Theodoxus ( Calvertia ) brusinai ( Pavlović, 1931 ) would be secondary homonym of Theodoxus ( Ninnia ) brusinai ( Andrusov, 1909 ) . However, Ninnia Brusina, 1902 was meanwhile elevated to genus level based on differences of shell and ontogeny ( Bandel 2001 , p. 128). A replacement name is thus not necessary at present. Another Theodoxus species formally constituting a secondary homonym is Theodoxus ( Calvertia ) depressus Taner, 1974 . The species-group name " depressus " in combination with Theodoxus is already in use for a taxon from the Pleistocene of Kos Island, introduced as " Neritina Fuchsi var. depressa " by Magrograssi (1928) and recombined as Theodoxus doricus depressus ( Magrograssi, 1928 ) by Willmann (1981 , p. 149). Wesselingh et al. (2008 , p. 865), however, consider the species described by Taner (1974) as junior synonym of Theodoxus bukowskii ( Oppenheim, 1919 ) , thus we refrain from introducing a replacement name.