Four new species of the grass feeding leafhopper genus Nicolaus Lindberg (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent Author Viraktamath, C. A. Author Webb, M. D. text Zootaxa 2014 3784 5 528 538 journal article 46085 10.11646/zootaxa.3784.5.2 9198978a-c098-4cba-94ef-97d747895d6a 1175-5326 228652 209DF328-B6EE-45DC-88E2-412C5E7D514F Key to species of Nicolaus Lindberg from the Indian subcontinent 1. Vertex of head with anterior submarginal brown stripe between eyes ( Figs 1, 3 , 9, 21)............................... 2 - Vertex of head without submarginal brown stripe between eyes, may have brown patches ( Figs 5 , 41 )... N. cornutus sp. nov. 2. Aedeagal shaft anteriorly sclerotized medially ( Figs 35 , 56 ); apophysis of style slender, without toothed pigmented border ( Figs 34 , 53 )......................................................................................... 3 - Aedeagal shaft membranous ( Figs 17 , 25 ); apophysis of style stouter with toothed pigmented border ( Figs15 , 23 )......... 4 3. Preatrial processes of aedeagus enlarged and strongly serrated subapically on ventral margin ( Fig. 56 )... N. serratus sp. nov. - Preatrial processes of aedeagus expanded apically with pair of prominent denticles on ventral margin ( Figs 36, 37 ).......................................................................................... N. bihamatus Xing & Li 4. Preatrial processes of aedeagus with pair of prominent denticles on ventral margin ( Figs 24, 25 )...... N. bidentatus sp. nov. - Preatrial processes of aedeagus without prominent denticles on ventral margin ( Figs 17, 18 )........... N. abuensis sp. nov.