Phylogenetic relationships in Coryphantha and implications on Pelecyphora and Escobaria (Cacteae, Cactoideae, Cactaceae) Author Sanchez, Daniel CONACYT-Laboratorio Nacional de Identificacion y Caracterizacion Vegetal, Departamento de Botanica y Zoologia, Centro Universitario de Ciencias Biologicas y Agropecuarias, Universidad de Guadalajara, Zapopan, Jalisco, C. P. 45220, Mexico & Herbario Luz Maria Villarreal de Puga, Departamento de Botanica y Zoologia, Centro Universitario de Ciencias Biologicas y Agropecuarias, Universidad de Guadalajara, Zapopan, Jalisco, C. P. 45220, Mexico Author Vazquez-Benitez, Balbina Coleccion de Plantas Suculentas, Facultad de Estudios Superiores Zaragoza, Campus II, Universidad Nacional Autonoma de Mexico, C. P. 15000, CDMX, Mexico Author Vazquez-Sanchez, Monserrat Programa de Posgrado en Botanica, Colegio de Postgraduados. Carretera Mexico-Texcoco Km 36.5, Montecillo, Texcoco, Estado de Mexico, 56230, Mexico vazquez.monserrat@colpos.mx Author Aquino, David Jardin Botanico, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Coyoacan, 04510, CDMX, Mexico Author Arias, Salvador https://orcid.org/0000-0002-7674-7050 Jardin Botanico, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Coyoacan, 04510, CDMX, Mexico sarias@ib.unam.mx text PhytoKeys 2022 2022-01-21 188 115 165 http://dx.doi.org/10.3897/phytokeys.188.75739 journal article http://dx.doi.org/10.3897/phytokeys.188.75739 1314-2003-188-115 86E2956EE98B5D6A9103EC1CE5B30E8F Pelecyphora missouriensis (Sweet) D.Aquino & Dan. Sanchez comb. nov. Mammillaria missouriensis ≡ Mammillaria missouriensis Sweet, Hort. Brit.: 171. 1826. Cactus missouriensis (Sweet) Kuntze, Revis. Gen. Pl. 1: 259. 1891. Mammillaria missouriensis Sweet ex K.Schum., Gesamtbeschr. Kakt.: 498. 1898. Coryphantha missouriensis (Sweet) Britton & Rose, Ill. Fl. N. U.S. 2: 570. 1913. Neobesseya missouriensis (Sweet) Britton & Rose, Cactaceae 4: 53. 1923. Neomammillaria missouriensis (Sweet) Britton & Rose ex Rydb., Fl. Plains N. Amer. 561. 1932. Escobaria missouriensis (Sweet) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type: United States, North Dakota, Burleigh County, 3 mi [4.8 km] W of Baldwin turnoff, 1.3 m [2.09 km] E of Missouri River, Jun 1970, L. Mitich s.n. (neotype, designated by Mitich & Benson, Cact. Succ. J. (Los Angeles): 8. 1977: POM [317949]; isoneotype NDA). Mammillaria similis = Mammillaria similis Engelm. & A.Gray, Boston J. Nat. Hist. 5: 246. 1845. Echinocactus similis (Engelm.) Poselg., Allg. Gartenzeitung 21: 107. 1853. Cactus missouriensis var. similis (Engelm.) J.M.Coult. in Contr. U.S. Natl. Herb. 3: 111. 1894. Cactus similis (Engelm.) Small, Fl. S.E. U.S.: 812. 1903. Coryphantha similis (Engelm.) Britton & Rose, Ill. Fl. N. U.S.: 571. 1913. Neobesseya similis (Engelm.) Britton & Rose, Cactaceae 4: 52. 1923. Escobaria missouriensis var. similis (Engelm.) N.P.Taylor, Kakteen And. Sukk. 34: 184. 1983. Neobesseya missouriensis subsp. similis (Engelm.) Doweld, Sukkulenty 3: 37. 2000. Type: United States, Sandstone rocks, near Industry (not preserved). Mammillaria nuttallii = Mammillaria nuttallii Engelm., Mem. Amer. Acad. Arts n.s., 4: 49. 1849. Coryphantha nuttallii Engelm. ex C.F. Foerst ., Handb. Cacteenk.: 407. 1885. Mammillaria missouriensis var. nuttallii (Engelm.) Schelle, Handb. Kakteenkult.: 241. 1907. Neobesseya nuttallii Boed., Mammill.-Vergl.-Schluessel: 15. 1933. Neobesseya nuttallii (Engelm.) Borg, Cacti: 303. 1937. Type. United States, South Dakota, Ft. Pierre, on the Upper Missouri, 1847, F. V. Hayden s.n. (lectotype, designated by Benson, Cacti U. S. Canada: 964. 1982: MO [899104 image!, 899105 image!, two sheets]). Mammillaria similis robustior = Mammillaria similis var. robustior Engelm., Boston J. Nat. Hist. 6: 200. 1850. Mammillaria nuttallii var. robustior (Engelm.) Engelm. & J.M.Bigelow, Pacif. Railr. Rep. 4: 28. 1856. Mammillaria missouriensis var. robustior (Engelm.) S.Watson, Bibl. Index N. Amer. Bot.: 440. 1878. Cactus missouriensis var. robustior (Engelm.) J.M.Coult., Contr. U.S. Natl. Herb. 3: 111. 1894. Coryphantha missouriensis var. robustior (Engelm.) L.D.Benson, Cact. Succ. J. (Los Angeles) 41: 190. 1969. Escobaria missouriensis var. robustior (Engelm.) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type: United States, Texas, Piedernales [ Perdenales ] [River, Texas], May 1846, F. Lindheimer s.n . (lectotype, designated by Benson, Cact. Succ. J. (Los Angeles): 190. 1969: MO [2017430 image!]). Mammillaria similis caespitosa = Mammillaria similis var. caespitosa Engelm., Boston J. Nat. Hist. 6: 200. 1850. Mammillaria nuttallii var. caespitosa Engelm., Proc. Amer. Acad. Arts 3: 265. 1856. Mammillaria missouriensis var. caespitosa (Engelm.) S.Watson, Smithsonian Misc. Collect. 258: 403. 1878. Mammillaria wissmannii Hildm. ex K.Schum., Gesamtbeschr. Kakt.: 498. 1898. nom. nov. Neobesseya wissmannii (Hildm. ex K.Schum.) Britton & Rose, Cactaceae 4: 52. 1923. Coryphantha wissmannii (Hildm. ex K.Schum.) A.Berger, Kakteen: 278. 1929. Coryphantha missouriensis var. caespitosa (Engelm.) L.D.Benson, Cact. Succ. J. (Los Angeles) 41: 189. 1969. Escobaria missouriensis var. caespitosa (Engelm.) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type: United States, Cult. In hort. Goebels , St Louis from Texas near Industry, May 1846, F. Lindheimer s.n . (lectotype, designated by Benson, Cact. Succ. J. (Los Angeles): 190. 1969: MO). Mammillaria nuttallii borealis = Mammillaria nuttallii var. borealis Engelm., Proc. Amer. Acad. Arts 3: 264. 1856. Type: United States, on the Upper Missouri. Not preserved. Mammillaria notesteinii = Mammillaria notesteinii Britton, Bull. Torrey Bot. Club 18: 367. 1891. Cactus notesteinii (Britton) Rydb., Mem. New York Bot. Gard. 1: 272. 1900. Neobesseya notesteinii (Britton) Britton & Rose, Cactaceae 4: 53. 1923. Type: United States, Deer Londge, Mont., 01 Jun 1891, F. N. Notestein s.n . (lectotype, designated here: NY [385874 image, four sheets!]; isolectotype: US [1821122 image!]). Coryphantha marstonii = Coryphantha marstonii Clover, Bull. Torrey Bot. Club 65: 412. 1938. Coryphantha missouriensis var. marstonii (Clover) L.D.Benson, Cacti Ariz. ed. 3: 26. 1969. Escobaria missouriensis var. marstonii (Clover) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type. United States, Utah, Kane County, east side of Buckskin Mountains, 5200 ft [1584 m], 08 Aug 1953, L. D. Benson & R. Benson 15205 (neotype, designated by Benson, Cacti Ariz. ed. 3: 26. 1969: POM [285320, 296309, two sheets]). Escobaria missouriensis navajoensis = Escobaria missouriensis subsp. navajoensis Hochstaetter , Succulenta (Netherlands) 75: 257. 1996. Type. United States, Arizona, Navajoa, 1600-1800, F. Hochstaetter 1000 (holotype: HBG).