To the revision of the genus Thionia Stål (Hemiptera, Fulgoroidea, Issidae), with description of new genera and new subtribe Author Gnezdilov, Dimir M. text Zootaxa 2018 2018-06-14 4434 1 158 170 journal article 29889 10.11646/zootaxa.4434.1.10 06e27461-4a99-402f-ae42-eb81da865394 1175-5326 1290884 B22116EA-DF00-4C6D-93FB-D40427809537 Fowlerium gen. n. Type species: Thionia naso Fowler, 1905 . Diagnosis. Metope elongate, with distinct median and sublateral carinae joined in one point on its upper margin ( Doering 1938, Pl. 52: 8, 10 ). Coryphe elongate, with distinct median carina running from its anterior margin to its middle; anterior margin acutely angulately convex ( Doering 1938, Pl. 51: 2, 5 ). Second and third segments of rostrum are almost equal in length; third segment narrowing apically ( Fig. 18 ). Fore wings elongate, surpassing abdomen, with narrow hypocostal plate. Forewing vein sequence: R 2, furcating near to basal cell; M 2, furcating before wing middle; CuΑ 1. Clavus long, open—Pcu+Α1 runs into apex of clavus. Pcu fusing Α1 after middle of clavus. Hind wings 3-lobed, with deep cubital and anal clefts and without coupling lobe on its anterior margin. CuΑ2 and CuP not fused apically, but connected by several transverse veins ( Figs 12, 13 ). Pcu and Α1.1 not fused medially, free. Hindwing sequence: R 2 r-m 1 M 1 m-cua 1 CuΑ 2 cua-cup 2-5 CuP 1 cup-pcu 1 Pcu 3 pcu-a1 1 Α1 2 Α2 2–4. Hind tibia with 2 lateral spines. First metatarsomere with a whole row of spines arranged in arc. Ventral aedeagal hooks slightly furcating subapically ( Doering, 1938, Pl. 55: 1, 7, 8 ). Ovipositor with rounded gonoplacs. Hind margin of VII sternum with large semicircular process. Etymology. Genus named in honor of famous British entomologist William Weeks Fowler ( 1849–1923 ) who describing Thionia naso had doubts on its generic position already ( Fowler 1905: 125 ). Neuter in gender. Composition. In addition to type species Thionia producta Van Duzee, 1908 and T. acuta Doering, 1941 also belong to this genus.