A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae) Author Sabella, Giorgio Dipartimento di Scienze Biologiche, Geologiche ed Ambientali dell’Università - sezione Biologia Animale, via Author Cuccodoro, Giulio Muséum d’histoire naturelle, CP 6434, CH- 1211 Genève 6, Switzerland. Author Kurbatov, Sergey A. Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail: text Revue suisse de Zoologie 2020 2020-06-04 127 1 129 156 http://dx.doi.org/10.35929/rsz.0013 journal article 117406 10.35929/RSZ.0013 8458d5d1-44ea-4c33-b453-b746ddaa2534 0035-418 5743391 Achilia delamarei Jeannel, 1962 (stat. nov.) Figs 2 , 13 , 23, 30 , 42 , 55-58 , 83 Achilia grandiceps delamarei Jeannel, 1962: 421 , 422. Type material ( 4 ex. ): SOUTHERN ARGENTINA : Rio Negro prov. : MNHN ; 1 ♂ ( lectotype , here designated); labels verbatim “Lectotype / Nahuel Huapi, B31, Cl. Delam. / grandiceps (handwritten by Jeannel) / Achilia delamarei ♂, Sabella, Cuccodoro & Kurbatov det. 2019”. – MNHN ; 1 ♀ ( paralectotype , here designated); same data as lectotype. – MNHN ; 2 ♀ ( paralectotypes , here designated); labels verbatim “Paralectotype / Parc Nat. Nahuel Huapi, B27 / Achilia delamarei ♀, Sabella, Cuccodoro & Kurbatov det. 2019” . Additional material examined ( 141 ex. ): SOUTHERN AND CENTRAL CHILE : Región Aysén : Aysén prov .: MHNG ; 1 ♀ ; 30 km N Puyuhuapi ; 100 m ; 29.I.1985 ; site 107, sifted moss on logs ; S. & J. Peck . – Región Los Lagos : Palena prov .: MHNG ; 20 ♂ and 58 ♀ ; 4 km NW Chaitén ; 10 m ; 30.I.1985 ; mixed forest litter , sooty fungus , berlese , S. & J. Peck . – FMNH ( FMHD #85-991 , P #85-108 ); 2 ♂ and 5 ♀ ; same data; S. & J. Peck . – PCTS ; 12 ♀ ; Homopirén , 41° 87’S 72° 36’W; 17.XII.2013 ; forest layer . – MHNG ; 1 ♂ and 1 ♀ ; same data. – Osorno prov. : UNHC ; 1 ♂ and 1 ♀ ; Puyehue National Park , 4.1 km E Anticura ; 430 m ; 19-26. XII.1982 ; trap site 662, valdivian rainforest ; A. Newton & M. Thayer . – FMNH ; 8 ♀ ; same data . – FMNH ( FMHD# 97-5 ); 1 ♂ ; Puyehue National Park , 4 km E Anticura ; 40° 39.73’S 72° 08.10’W ; 460 m ; 30.I.1997 ; site, 985-3 valdivian rainforest w/large Saxegothea , flight intercept trap ; A. Newton & M. Thayer . – MHNG ; 2 ♂ ; Puyehue National Park , Aguas Calientes ; 500 m ; 20.XII.1984 - 08.II.1985 ; Nothofagus forest; S. & J. Peck . – PCTS ; 2 ♂ and 5 ♀ ; Aguas Calientes , 40° 74’S 72° 30’W; 13.XII.2013 ; car net . – PCTS ; 3 ♂ and 7 ♀ ; Aguas Calientes , 40° 74’S 72°27’W; 14.XII.2013 ; litter layer . – FMNH ( FMHD #2002-083 ); 1 ♀ ; Vicente Perez Rosales National Park , SW slope Volcàn Osorno , road to Ref. La Picada ; 41° 01.05’S 72° 32.90’W ; 430 m ; 16.XII.2002 ; site 1068, Nothofagus dombeyi w/conifers, berlese, leaf & log litter; A. Newton , A. Solodovnikov & M. Chani . – Región Araucanía : Cautín prov.: MHNG ; 1 ♂ and 9 ♀ ; Huerquehue National Park ; 800-900 m ; 22-24.XII.1980 ; site 16a, forest litter ; D. Agosti & D. Burckhardt . Description: Body 1.55-1.75 mm long, dark brown with reddish elytra, sometimes darker basally and apically, and along sutural stria; antennae and legs reddish or reddish-brown; palpi yellowish. Pronotum with disc slightly more convex than in A. grandiceps ; posterior portion of lateral margins subparallel. First abdominal tergite with diverging basal striae extending to about one-fourth of paratergal length, and separated at base by about one-third of tergal width. Male : Head as in Figs 55-58 , similar to that of A. grandiceps except: the two quadrangular protuberances wider than long, and separated by U-shaped median notch narrower at base than that for A. grandiceps ; lateral arms of notch densely pubescent apically; anterior portion of frons very deeply excavated; median apophysis originating from very broad median sub-basal clypeal hump, tip in dorsal view lozenge-shaped with pubescent sides; median sub-basal clypeal hump with sides sharply margined and bearing only some short setae. Also in this species the head morphology of all the male specimens does not show substantial differences. Antennae ( Fig. 13 ) with scape slightly longer than wide and slightly enlarged on lateral margin; pedicel distinctly longer than wide; antennomere III longer than wide; antennomere IV as wide as long; antennomere V longer than wide; antennomeres VI-VIII wider than long; antennomere IX wider than VIII and wider than long; antennomere X wider than long; antennomere XI elongate, and longer than VII-X combined. Metaventrite with distal half pubescent, raised at middle for two thirds of its distal portion, this surface entirely divided by broad medial sulcus. Protrochanters ( Fig. 23 ) bearing one long seta; mesotrochanters ( Fig. 30 ) with ventral margin forming spine on basal third; mesofemora ( Fig. 30 ) with ventral margin covered by broad, short and thick setae on basal third, these setae longer than in A. grandiceps ; mesotibiae with medial margin forming short and rounded apical spine, always with recurved setae ( Fig. 42 ); apical margin forming second tiny spine at middle. Aedeagus ( Fig. 2 ) 0.32-0.33 mm long, similar to that of A. grandiceps except lateral sclerites thicker, distinctly pointed and apically recurved, and medial sclerites with two-thirds of distal portion frayed and bearing numerous long and thin spines. Female : Similar to male except body sometimes paler than male, head not modified with frons depressed behind frontal sulcus which is well-impressed, and anterior margin of frontal lobe pointed at middle. Collecting data: Collected from December to March in different types of forests, at elevations ranging from 10 m to 900 m . Most specimens came from sifted samples of leaf and log litter, and some males have also been collected by flight intercept traps and car netting. Distribution: Achilia delamarei is distributed in southern Argentina and southern and central Chile ( Fig. 83 : red squares) ranging from Aysén province to Cautín province. Comments: Jeannel (1962: 422) in the original description mentions 1 male and 3 females collected by Cl. Delamare in March 1959 in the province of Rio Negro (Argentina) in the Nahuel Huapi reserve (about 41° latitude S) at an altitude of about 1000 meters, and claimed (l. c.: 421) that the type of this taxon is in the MNHN collections. In the general collection of Chilean Pselaphinae at the MNHN we found 1 male and 3 females labeled as being from Nahuel Huapi , with only the male bearing a label “ grandiceps (handwritten by Jeannel)”, and none bearing red “ Type ” labels. For this reason, we designate here the only male of this series as the lectotype and the 3 females as paralectotypes . The lectotype of A. delamarei has all antennomeres slightly longer that those of the other males examined, in particular the pedicel is two times longer than wide, while in the other males it is typically one and a half times longer than wide. For the distinctive characters that separate A. delamarei and A. grandiceps see the “ Comments ” section under A. grandiceps .