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<mods:title id="3FDE0D4C0BCAB3C9C3F48C6C847216BA">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<emphasis id="D6E0C41851581835FF0EFED2EE601D2F" bold="true" box="[151,344,333,360]" italics="true" pageId="99" pageNumber="100">Luciola parvula</emphasis>
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<emphasis id="D6E0C41851581835FF0EFE0EEE051DEE" box="[151,317,401,425]" italics="true" pageId="99" pageNumber="100">Luciola parvula</emphasis>
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<bibRefCitation id="800565FB51581835FEDFFE0EED791DEE" author="Kiesenwetter" box="[326,577,401,426]" firstAuthor="Kiesenwetter" pageId="99" pageNumber="100" pagination="241 - 288" refId="ref118656" refString="Kiesenwetter, H. (1874) Die Malacodermen Japans nach den Ergebnisse der Sammlungen des Herrn L. Lewis wahrend der Jahre 1869 - 1871. Deutsche Entomologische Zeitschrift, 18, 241 - 288. https: // doi. org / 10.1002 / mmnd. 18740180302" type="journal article" year="1874">Kiesenwetter 1874: 241</bibRefCitation>
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<treatmentCitation id="65353E1B51581835FDD7FE0EEC2D1DED" author="Gorham, H. S." box="[590,789,401,426]" page="410" pageId="99" pageNumber="100" year="1883">
<bibRefCitation id="800565FB51581835FDD7FE0EEC2D1DED" author="Gorham" box="[590,789,401,426]" firstAuthor="Gorham" pageId="99" pageNumber="100" pagination="393 - 411" refId="ref117533" refString="Gorham, H. S. (1883) Review of the genera and species of Malacoderm Coleoptera of the Japanese fauna. Transactions of the Entomological Society of London, 1883, 393 - 411. https: // doi. org / 10.1111 / j. 1365 - 2311.1883. tb 02955. x" type="journal article" year="1883">Gorham 1883: 410</bibRefCitation>
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<treatmentCitation id="65353E1B51581835FC81FE0EECF21DED" author="Olivier, E." box="[792,970,401,426]" page="85" pageId="99" pageNumber="100" year="1902">
<bibRefCitation id="800565FB51581835FC81FE0EECF21DED" author="Olivier" box="[792,970,401,426]" firstAuthor="Olivier" pageId="99" pageNumber="100" pagination="69 - 88" refId="ref120443" refString="Olivier, E. (1902) Catalogue des especes de ' Luciola' et genres voisins decrits jusqu'a ce jour. Revue Scientifique du Bourbonnais et du centre de la France, 15, 69 - 88." type="journal article" year="1902">Olivier 1902: 85</bibRefCitation>
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; 
<treatmentCitation id="65353E1B51581835FC41FE0EEB0F1DED" author="Olivier, E." box="[984,1079,401,426]" page="53" pageId="99" pageNumber="100" year="1907">
<bibRefCitation id="800565FB51581835FC41FE0EEB0F1DED" author="Olivier" box="[984,1079,401,426]" firstAuthor="Olivier" pageId="99" pageNumber="100" pagination="1 - 74" refId="ref120599" refString="Olivier, E. (1907) Coleoptera, Lampyridae. Genera Insectorum, 53, 1 - 74. [Luciolinae, 48 - 56]." type="journal article" year="1907">1907: 53</bibRefCitation>
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; 
<treatmentCitation id="65353E1B51581835FBDCFE0EEB881DED" author="Olivier, E." box="[1093,1200,401,426]" page="45" pageId="99" pageNumber="100" year="1910">
<bibRefCitation id="800565FB51581835FBDCFE0EEB881DED" author="Olivier" box="[1093,1200,401,426]" firstAuthor="Olivier" pageId="99" pageNumber="100" pagination="40 - 48" refId="ref120761" refString="Olivier, E. (1910 a) Pars 9. Lampyridae. In: Coleopterorum Catalogus (Junk, Schenkling). Junk, Berlin, pp. 1 - 68. [Luciolinae, 40 - 48]" type="book chapter" year="1910" yearSuffix="a">1910a: 45</bibRefCitation>
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<treatmentCitation id="65353E1B51581835FB24FE0DEAA11DED" author="Matsumura, M." box="[1213,1433,401,426]" page="60" pageId="99" pageNumber="100" year="1928">
<bibRefCitation id="800565FB51581835FB24FE0DEAA11DED" author="Matsumura" box="[1213,1433,401,426]" firstAuthor="Matsumura" pageId="99" pageNumber="100" pagination="39 - 70" refId="ref119255" refString="Matsumura, M. (1928) The Fireflies. In: Using interesting insects as teaching tool. Tokyodo Shoten, Tokyo, pp. 39 - 70. [in Japanese]" type="book chapter" year="1928">Matsumura 1928: 60</bibRefCitation>
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<paragraph id="E42B180A51581835FF5EFE30EAA51E42" blockId="99.[151,1437,401,640]" pageId="99" pageNumber="100">
<bibRefCitation id="800565FB51581835FF5EFE30EE411D8F" author="Okada" box="[199,377,431,456]" firstAuthor="Okada" pageId="99" pageNumber="100" pagination="130 - 149" refId="ref120083" refString="Okada, Y. (1931) Notes on the scientific names of the Japanese fireflies. Zoological Magazine Tokyo, 43, 130 - 149. [in Japanese]" type="journal article" year="1931">Okada 1931: 146</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FE1BFE30ED1F1D8F" author="Kanda" box="[386,551,431,456]" firstAuthor="Kanda" pageId="99" pageNumber="100" refId="ref118530" refString="Kanda, S. (1935) Fireflies (Hotaru). The Association of Bioluminescence Research of Japan, Tokyo, 496 pp. [In Japanese]" type="book" year="1935">Kanda 1935: 81</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FDA8FE30EC691D8F" author="Kawashima" box="[561,849,431,456]" etAl="et al." firstAuthor="Kawashima" pageId="99" pageNumber="100" pagination="241 - 261" refId="ref118611" refString="Kawashima, I., Suzuki, H. &amp; Sato, M. (2003) A Check-List of Japanese Fireflies (Coleoptera, Lampyridae and Rhagophthalmidae). Japanese Journal of Systematic Entomology, 9 (2), 241 - 261." type="journal article" year="2003">
Kawashima 
<emphasis id="D6E0C41851581835FD36FE2EEDE31D8F" box="[687,731,432,456]" italics="true" pageId="99" pageNumber="100">et al</emphasis>
. 2003: 248
</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FCC3FE2FECE01D8F" author="Fu" box="[858,984,432,456]" firstAuthor="Fu" pageId="99" pageNumber="100" refId="ref116462" refString="Fu, X. H. (2014) An illustrated handbook of Chinese fireflies. The Commercial Press, Beijing, 167 pp. [in Chinese]" type="book" year="2014">Fu 2014: 37</bibRefCitation>
. 
<taxonomicName id="2394638951581835FF0EFE50ED901DA0" ID-CoL="5H69" authority="Paruula Kiesenwetter. Matsumura 1928: 83" authorityName="Paruula Kiesenwetter. Matsumura" authorityPageNumber="83" authorityYear="1928" box="[151,680,462,487]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851581835FF0EFE50EE061DA0" box="[151,318,463,487]" italics="true" pageId="99" pageNumber="100">Luciola Paruula</emphasis>
Kiesenwetter. 
<bibRefCitation id="800565FB51581835FE4CFE50ED901DA0" author="Matsumura" box="[469,680,463,487]" firstAuthor="Matsumura" pageId="99" pageNumber="100" pagination="39 - 70" refId="ref119255" refString="Matsumura, M. (1928) The Fireflies. In: Using interesting insects as teaching tool. Tokyodo Shoten, Tokyo, pp. 39 - 70. [in Japanese]" type="book chapter" year="1928">Matsumura 1928: 83</bibRefCitation>
</taxonomicName>
(misspelling). 
<taxonomicName id="2394638951581835FF0EFE71EEE41E41" authority="(Kiesenwetter)" baseAuthorityName="Kiesenwetter" box="[151,476,493,518]" class="Insecta" family="Lampyridae" genus="Hotaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="D6E0C41851581835FF0EFE71EE041E42" box="[151,316,493,517]" italics="true" pageId="99" pageNumber="100">Hotaria parvula</emphasis>
(Kiesenwetter)
</taxonomicName>
. 
<bibRefCitation id="800565FB51581835FE7DFE71EDA81E42" author="Yuasu" box="[484,656,493,517]" firstAuthor="Yuasu" pageId="99" pageNumber="100" pagination="107 - 109" refId="ref122645" refString="Yuasu, H. (1937) Description of a new genus of the family Lampyridae. Kontyu, XI, 107 - 109." type="journal article" year="1937">Yuasu 1937: 107</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FD03FE71EC491E42" author="McDermott" box="[666,881,493,518]" firstAuthor="McDermott" pageId="99" pageNumber="100" pagination="1 - 72" refId="ref119352" refString="McDermott, F. A. (1964) The taxonomy of the Lampyridae. Transactions of the American Entomological Society, 90, 1 - 72." type="journal article" year="1964">McDermott 1964: 44</bibRefCitation>
; 
<bibRefCitation id="800565FB51581835FCE5FE72ECDB1E42" author="McDermott" box="[892,995,493,518]" firstAuthor="McDermott" pageId="99" pageNumber="100" pagination="1 - 149" refId="ref119380" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.), Coleopterorum Catalogus Supplementa. Pars 9. Editio Secunda. W. Junk, S'Gravenhage, pp. 1 - 149." type="book chapter" year="1966">1966: 114</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FC74FE72EB9F1E42" author="Nakane" box="[1005,1191,493,518]" firstAuthor="Nakane" pageId="99" pageNumber="100" pagination="118 - 119" refId="ref119774" refString="Nakane, T. (1960) Coloured illustrations of the insects of Japan. Coleoptera. Kinki Coleopterological Society, Hoikusha, Osaka, 385 pp. [Lampyridae 118 - 119]" type="book chapter" year="1960">Nakane 1960: 119</bibRefCitation>
; 
<bibRefCitation id="800565FB51581835FB28FE72EA351E41" author="Nakane" box="[1201,1293,493,518]" firstAuthor="Nakane" pageId="99" pageNumber="100" pagination="110 - 113" refId="ref119853" refString="Nakane, T. &amp; Ohba, N. (1981) Hotaru no Kansatsu to Shiiku. Observations and rearing of fireflies. Nyu Saiensusha, Tokyo, 122 pp. [in Japanese; (Lampyridae 110 - 113]" type="book chapter" year="1981">1981: 96</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FA8EFE72EAA51E42" author="Nakane" box="[1303,1437,493,518]" etAl="et al." firstAuthor="Nakane" pageId="99" pageNumber="100" pagination="169 - 170" refId="ref119895" refString="Nakane, T., Ohbayashi, K., Nomura, S. &amp; Kurosawa, Y. (1965) Iconographia Insectorum Japonicorum Colore naturali edita. Vol. II. Coleoptera. Hokuryukan, Tokyo, 443 pp. [Lampyridae 169 - 170, pl. 85]" type="book chapter" year="1965">
Nakane 
<emphasis id="D6E0C41851581835FAF3FE71EAA51E42" box="[1386,1437,493,517]" italics="true" pageId="99" pageNumber="100">et al.</emphasis>
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151581835FF5EFD94EDB11EC7" pageId="99" pageNumber="100" type="description">
<paragraph id="E42B180A51581835FF5EFD94EDB11EC7" blockId="99.[151,1437,401,640]" pageId="99" pageNumber="100">
1965: 170. 
<bibRefCitation id="800565FB51581835FEA3FD94ED741E63" author="Nakane &amp; Ohba" box="[314,588,523,548]" firstAuthor="Nakane" pageId="99" pageNumber="100" pagination="110 - 113" refId="ref119853" refString="Nakane, T. &amp; Ohba, N. (1981) Hotaru no Kansatsu to Shiiku. Observations and rearing of fireflies. Nyu Saiensusha, Tokyo, 122 pp. [in Japanese; (Lampyridae 110 - 113]" type="book chapter" year="1981">Nakane &amp; Ohba 1981: 111</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FDCFFD94EDDA1E63" author="Ohba" box="[598,738,523,548]" firstAuthor="Ohba" pageId="99" pageNumber="100" pagination="1 - 62" refId="ref120015" refString="Ohba, N. (1983) Studies on the communication system of Japanese fireflies. Science Report of the Yokosuka City Museum, 30, 1 - 62." type="journal article" year="1983">Ohba 1983: 8</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FD75FD94ECC31E63" author="Hayashi &amp; Suzuki" box="[748,1019,523,548]" firstAuthor="Hayashi" pageId="99" pageNumber="100" pagination="3 - 10" refId="ref117886" refString="Hayashi, F. &amp; Suzuki, H. (2003) Fireflies with or without prespermatophores: Evolutionary origins and life-history consequences. Entomological Science, 6, 3 - 10. https: // doi. org / 10.1046 / j. 1343 - 8786.2003.00003. x" type="journal article" year="2003">Hayashi &amp; Suzuki 2003: 5</bibRefCitation>
. 
<taxonomicName id="2394638951581835FF0EFDB4ED791E04" authority="(Kiesenwetter)" baseAuthorityName="Kiesenwetter" box="[151,577,554,579]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="parvula" subGenus="Hotaria">
<emphasis id="D6E0C41851581835FF0EFDB4EFDC1E04" box="[151,228,555,579]" italics="true" pageId="99" pageNumber="100">Luciola</emphasis>
(
<emphasis id="D6E0C41851581835FF6AFDB4EE7B1E05" box="[243,323,555,578]" italics="true" pageId="99" pageNumber="100">Hotaria</emphasis>
) 
<emphasis id="D6E0C41851581835FECBFDB4EE991E04" box="[338,417,555,579]" italics="true" pageId="99" pageNumber="100">parvula</emphasis>
(Kiesenwetter)
</taxonomicName>
. 
<bibRefCitation id="800565FB51581835FDD2FDB4EC171E04" author="McDermott" box="[587,815,554,579]" firstAuthor="McDermott" pageId="99" pageNumber="100" pagination="1 - 149" refId="ref119380" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.), Coleopterorum Catalogus Supplementa. Pars 9. Editio Secunda. W. Junk, S'Gravenhage, pp. 1 - 149." type="book chapter" year="1966">McDermott 1966: 114</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FCA0FDB5EB2E1E04" author="Ballantyne" box="[825,1046,554,579]" firstAuthor="Ballantyne" pageId="99" pageNumber="100" pagination="103 - 139" refId="ref113808" refString="Ballantyne, L. A. (1968) Revisional Studies of Australian and Indomalayan Luciolini (Coleoptera, Lampyridae, Luciolinae). University of Queensland Papers, Department of Entomology, University of Queensland Press, II (6), 103 - 139." type="journal article" year="1968">Ballantyne 1968: 124</bibRefCitation>
. 
<taxonomicName id="2394638951581835FF0EFDD6ED271E25" authority="Nawa 1921: 256" authorityName="Nawa" authorityPageNumber="256" authorityYear="1921" box="[151,543,585,610]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="ibukiyamana">
<emphasis id="D6E0C41851581835FF0EFDD6EE561E26" box="[151,366,585,609]" italics="true" pageId="99" pageNumber="100">Luciola ibukiyamana</emphasis>
<bibRefCitation id="800565FB51581835FEEDFDD5ED271E25" author="Nawa" box="[372,543,585,610]" firstAuthor="Nawa" pageId="99" pageNumber="100" pagination="256 - 260" refId="ref119986" refString="Nawa, U. (1921) Report on the Ibuki-firefly, Luciola ibukiyamana. Insect World, Gifu 25, 256 - 260. [in Japanese]" type="journal article" year="1921">Nawa 1921: 256</bibRefCitation>
</taxonomicName>
. 
<bibRefCitation id="800565FB51581835FDB1FDD5EC361E25" author="McDermott" box="[552,782,585,610]" firstAuthor="McDermott" pageId="99" pageNumber="100" pagination="1 - 149" refId="ref119380" refString="McDermott, F. A. (1966) Lampyridae. In: Steel, W. O. (Ed.), Coleopterorum Catalogus Supplementa. Pars 9. Editio Secunda. W. Junk, S'Gravenhage, pp. 1 - 149." type="book chapter" year="1966">McDermott 1966: 105</bibRefCitation>
. 
<bibRefCitation id="800565FB51581835FC8EFDD6EB0C1E26" author="Kawashima" box="[791,1076,585,610]" etAl="et al." firstAuthor="Kawashima" pageId="99" pageNumber="100" pagination="241 - 261" refId="ref118611" refString="Kawashima, I., Suzuki, H. &amp; Sato, M. (2003) A Check-List of Japanese Fireflies (Coleoptera, Lampyridae and Rhagophthalmidae). Japanese Journal of Systematic Entomology, 9 (2), 241 - 261." type="journal article" year="2003">
Kawashima 
<emphasis id="D6E0C41851581835FC0CFDD5ECFA1E26" box="[917,962,585,609]" italics="true" pageId="99" pageNumber="100">et al</emphasis>
. 2003: 248
</bibRefCitation>
(synonymy). 
<emphasis id="D6E0C41851581835FF0EFDF7EFF41EC7" bold="true" box="[151,204,616,640]" pageId="99" pageNumber="100">
<typeStatus id="3B2FA6A851581835FF0EFDF7EFF41EC7" box="[151,204,616,640]" pageId="99" pageNumber="100">Type</typeStatus>
</emphasis>
. 
<taxonomicName id="2394638951581835FF40FDF7EE441EC7" box="[217,380,616,640]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="D6E0C41851581835FF40FDF7EE441EC7" box="[217,380,616,640]" italics="true" pageId="99" pageNumber="100">Luciola parvula</emphasis>
</taxonomicName>
. Male. 
<emphasis id="D6E0C41851581835FE50FDF8ED181EC7" bold="true" box="[457,544,615,640]" pageId="99" pageNumber="100">
<collectingCountry id="9C83589A51581835FE50FDF8ED241EC7" box="[457,540,615,640]" name="Japan" pageId="99" pageNumber="100">JAPAN</collectingCountry>
.
</emphasis>
(NHML).
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151581835FF0EFD32ED591F57" pageId="99" pageNumber="100" type="diagnosis">
<paragraph id="E42B180A51581835FF0EFD32ED591F57" blockId="99.[151,1437,685,784]" pageId="99" pageNumber="100">
<emphasis id="D6E0C41851581835FF0EFD32EE321E8F" bold="true" box="[151,266,685,712]" pageId="99" pageNumber="100">Diagnosis</emphasis>
. A Japanese species with pale pronotum having an anteromedian dark marking, and dark brown to black elytra. Females are flightless with elytra covering or almost covering abdomen in repose, and no hind wings. 
<bibRefCitation id="800565FB51581835FAE6FD4CEFDA1F57" author="Fu" firstAuthor="Fu" pageId="99" pageNumber="100" refId="ref116462" refString="Fu, X. H. (2014) An illustrated handbook of Chinese fireflies. The Commercial Press, Beijing, 167 pp. [in Chinese]" type="book" year="2014">Fu (2014)</bibRefCitation>
recorded this species from 
<collectingCountry id="9C83589A51581835FD8EFD69ED661F57" box="[535,606,758,784]" name="China" pageId="99" pageNumber="100">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>