Revision of Poa L. (Poaceae, Pooideae, Poeae, Poinae) in Mexico: new records, re-evaluation of P. ruprechtii, and two new species, P. palmeri and P. wendtii
Author
Soreng, Robert J.
Department of Botany, National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA
Author
Peterson, Paul M.
Department of Botany, National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA
text
PhytoKeys
2012
2012-08-06
15
1
104
http://dx.doi.org/10.3897/phytokeys.15.3084
journal article
http://dx.doi.org/10.3897/phytokeys.15.3084
1314-2003-15-1
FF9AC356FFDAFB26FF95FFE3FFA1520C
576134
16a.
Poa pratensis agassizensis (B. Boivin & D.
Loeve
) Roy L. Taylor & MacBryde, Canad. J. Bot. 56(2): 193. 1978.
Fig. 17 A-C
Poa agassizensis
B. Boivin & D.
Loeve
, Naturaliste Canad. 87(6-7): 176-180, f. 1-2. 1960. Type: Canada, Manitoba, MacDonald Co, 6 Jun 1952,
B.Boivin,
Loeve
& Alex 9167
(holotype DAO!; isotype: US-2553819!).
Description.
Tufts sparse, some shoots clustered; pale green or bluish-gray-green; tillers intra- and extravaginal.
Culms
20-40(-50) cm tall.
Ligules
of lower culm and tiller leaves commonly glabrous abaxially; lades of cauline leaves flag leaf blades folded, with involute margins, moderately thick, moderately firm; sterile shoot blades usually less than 10 cm long, 0.8-2 mm wide, all folded with involute margins, sparsely pubescent adaxially.
Panicles
4-6(-8) cm long, erect or nodding, or loosely contracted or open, ovoid to narrowly pyramidal; rachis with 2-3(-5) branches per node; primary branches steeply ascending to ascending, smooth or sparsely to moderately densely scabrous; longest 1-2.5(-3) cm, with several spikelets per branch.
Spikelets
lanceolate, not bulbiferous; glumes unequal, glaucous or not; lower glumes 1(-3)-veined; upper glumes shorter than or nearly subequaling the lowest lemma; lemmas 2-3(-3.5) mm long, finely muriculate, intermediate veins glabrous; paleas scabrous, medially glabrous over the keels, intercostal region glabrous.
Anthers
frequently sterile. 2
n
= 41, 42, 43, 56.
Distribution.
This subspecies in North America is known from Canada and USA, and in Mexico from the states of Baja California and Coahuila.
Ecology.
The subspecies inhabits cool temperate prairies, meadows, and open coniferous forests.
Specimens examined.
Mexico.
Baja California:
Municipio Ensenada, Paraje El Rayo Ejido, 1000 m, 13 May 1997, A.Miranda 1153 (MEXU). Sierra San Pedro
Martir
, La Gurulla,
30°54'N
,
115°29'W
, 2050 m, 29 Jun 1982, R.Moran 30956 (MEXU, TAES).
Coahuila:
Sierra de Arteaga, Las Vigas, Canon de la Carbonera,
25°20'N
,
100°39'W
, 2100 m, 5 Jun 1987, J.A.Villarreal 3800 & M.A.Carranza (TAES). Municipio Arteaga, Ejido La Escondida,
25°23'30"N
,
100°33'15"W
, J.A.Garcia 49 (MEXU).
Discussion.
This form of
Poa pratensis
, originally described as a species indigenous to Canada and the United States (Boivin and
Loeve
1960) and now accepted as
Poa pratensis
subsp.
agassizensis
, is probably indigenous, but morphologically is questionably distinct from the Eurasian elements of
Poa pratensis
subsp.
angustifolia
, or xeric forms of cultivated
Poa pratensis
subsp.
pratensis
(
Soreng 2007
). As originally described it is a taxon with short, ovoid panicles, with small spikelets, and involute leaves. Such specimens are not uncommon among early herbarium collections across Canada and the USA. It approaches
Poa pratensis
subsp.
angustifolia
closely, but that subspecies of Eurasian origin has looser panicles, and is generally a lower polyploid (
Stoneberg-Holt 2004
). Separation of
Poa pratensis
subsp.
agassizensis
from
Poa pratensis
subsp.
pratensis
is
difficult
, and so far there is no molecular or cytological data to support it as a separate taxon. We recognize
Poa pratensis
subsp.
agassizensis
as a marginally distinct taxon, and site a few vouchers that best match this form.