<document id="664AC556D5ECA78EDD77DDCE9E270769" ID-DOI="10.17109/AZH.64.4.277.2018" ID-ISSN="2064-2474" ID-Zenodo-Dep="12626311" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638277157792" checkinUser="felipe" docAuthor="Nabozhenko, Maxim &amp; Ando, Kiyoshi" docDate="2018" docId="03BBAE26FFFFFFF0FEADA1D3548D3AB8" docLanguage="en" docName="ActaZoolAcadSciH.64.4.277-327.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 64 (4)" docSource="http://dx.doi.org/10.17109/azh.64.4.277.2018" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.8:ActaZoolAcadSciH.2015-2023.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-2023.journal_article" docStyleVersion="8" docTitle="Nipponohelops Masumoto, Ando" docType="treatment" docVersion="2" lastPageNumber="282" masterDocId="FF82D65EFFFAFFF5FFCCA16D56513801" masterDocTitle="Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region" masterLastPageNumber="327" masterPageNumber="277" pageNumber="282" updateTime="1719929912661" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="4C808EE5E30C378D0D6FE21776419DDB">Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region</mods:title>
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<mods:namePart id="36743F2302BE74BB1CA3E33F04C30870">Nabozhenko, Maxim</mods:namePart>
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<mods:namePart id="0563587F89B071DC59B979BBA9A43CDA">Ando, Kiyoshi</mods:namePart>
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<subSubSection id="C3084CBBFFFFFFF0FEADA1D355E238D9" box="[353,947,190,217]" pageId="5" pageNumber="282" type="nomenclature">
<paragraph id="8BAD1F30FFFFFFF0FEADA1D355E238D9" blockId="5.[353,947,190,217]" box="[353,947,190,217]" pageId="5" pageNumber="282">
<heading id="D0E5A85CFFFFFFF0FEADA1D355E238D9" box="[353,947,190,217]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="282" reason="2">
<taxonomicName id="4C1264B3FFFFFFF0FEADA1D354AD38D9" authority="Masumoto, Ando" authorityName="Masumoto, Ando" box="[353,764,190,217]" class="Insecta" family="Tenebrionidae" genus="Nipponohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis id="B966C322FFFFFFF0FEADA1D3544038D9" box="[353,529,190,216]" italics="true" pageId="5" pageNumber="282">Nipponohelops</emphasis>
Masumoto, Ando
</taxonomicName>
et 
<collectingRegion id="49D6D1D2FFFFFFF0FCEDA1D3553D38D9" box="[801,876,190,216]" country="Japan" name="Akita" pageId="5" pageNumber="282">Akita</collectingRegion>
, 2006
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3084CBBFFFFFFF0FF26A19A548D3AB8" pageId="5" pageNumber="282" type="materials_examined">
<paragraph id="8BAD1F30FFFFFFF0FF26A19A553E3928" blockId="5.[174,1126,246,326]" pageId="5" pageNumber="282">
<typeStatus id="54A9A192FFFFFFF0FF26A19A5771390D" box="[234,288,247,268]" pageId="5" pageNumber="282">Type</typeStatus>
species 
<emphasis id="B966C322FFFFFFF0FEB9A19B5434390A" box="[373,613,246,267]" italics="true" pageId="5" pageNumber="282">
<taxonomicName id="4C1264B3FFFFFFF0FEB9A19B5454390A" box="[373,517,246,267]" class="Insecta" family="Tenebrionidae" genus="Nipponohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="282" phylum="Arthropoda" rank="genus">Nipponohelops</taxonomicName>
ishikawai
</emphasis>
Masumoto, Ando et 
<collectingRegion id="49D6D1D2FFFFFFF0FC8DA19B552F390D" box="[833,894,246,268]" country="Japan" name="Akita" pageId="5" pageNumber="282">Akita</collectingRegion>
, 2006, by original designation. 
<bibRefCitation id="EF8362C1FFFFFFF0FEDEA079544A3929" author="MASUMOTO, K." box="[274,539,275,297]" pageId="5" pageNumber="282" pagination="33 - 38" refId="ref18990" refString="MASUMOTO, K., ANDO, Κ. &amp; AKITA, Κ. (2006): New or little known tenebrionid species from Japan (Part 5). A new species belonging to a new genus (Coleoptera: Tenebrionidae: Helopini). - Entomological Review of Japan 61: 33 - 38." type="journal article" year="2006">
Masumoto 
<emphasis id="B966C322FFFFFFF0FE47A07957E23929" box="[395,435,275,297]" italics="true" pageId="5" pageNumber="282">et al</emphasis>
., 2006: 33
</bibRefCitation>
; 
<bibRefCitation id="EF8362C1FFFFFFF0FDE9A07E553A3928" author="NABOZHENKO, M. V. &amp; LOBL, I." box="[549,875,275,297]" pageId="5" pageNumber="282" pagination="241 - 257" refId="ref19945" refString="NABOZHENKO, M. V. &amp; LOBL, I. (2008) Tribe Helopini. Pp. 241 - 257. In: LOBL, I. &amp; SMETANA, A. (eds): Catalogue of Palearctic Coleoptera. Vol. 5. Tenebrionoidea. Apollo Books, Stenstrup." type="book chapter" year="2008">Nabozhenko &amp; Löbl, 2008: 253</bibRefCitation>
.
</paragraph>
<paragraph id="8BAD1F30FFFFFFF0FF26A05D55AC3944" blockId="5.[174,1126,246,326]" box="[234,1021,304,326]" pageId="5" pageNumber="282">The detailed species description was given in the description of the genus.</paragraph>
<paragraph id="8BAD1F30FFFFFFF0FEEBA01355BF3999" blockId="5.[295,1006,382,409]" box="[295,1006,382,409]" pageId="5" pageNumber="282">
<emphasis id="B966C322FFFFFFF0FEEBA013541D3999" box="[295,588,382,408]" italics="true" pageId="5" pageNumber="282">
<taxonomicName id="4C1264B3FFFFFFF0FEEBA01357863999" box="[295,471,382,408]" class="Insecta" family="Tenebrionidae" genus="Nipponohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="282" phylum="Arthropoda" rank="genus">Nipponohelops</taxonomicName>
ishikawai
</emphasis>
Masumoto, Ando et 
<collectingRegion id="49D6D1D2FFFFFFF0FC90A01355F63999" box="[860,935,382,408]" country="Japan" name="Akita" pageId="5" pageNumber="282">Akita</collectingRegion>
, 2006
</paragraph>
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<heading id="D0E5A85CFFFFFFF0FD91A0CF54E639BD" box="[605,695,418,444]" centered="true" fontSize="11" level="3" pageId="5" pageNumber="282" reason="3">
(
<figureCitation id="132903B5FFFFFFF0FDAAA0CF54FD39BD" box="[614,684,418,444]" captionStart="Fig" captionStartId="4.[136,172,1505,1527]" captionTargetBox="[165,1065,201,1464]" captionTargetId="figure-19@4.[157,1069,189,1474]" captionTargetPageId="4" captionText="Fig. 1. Nipponohelops ishikawai: A = male, habitus, B = female, habitus, C = aedeagus, dorsal view, D = fore leg of male, E = middle leg of male, F = prothorax, ventral view, G = holotype" figureDoi="http://doi.org/10.5281/zenodo.12626315" httpUri="https://zenodo.org/record/12626315/files/figure.png" pageId="5" pageNumber="282">Fig. 1</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BAD1F30FFFFFFF0FF26A0B755F539EE" blockId="5.[173,1127,473,697]" box="[234,932,473,496]" pageId="5" pageNumber="282">
<bibRefCitation id="EF8362C1FFFFFFF0FF26A0B757A539EE" author="MASUMOTO, K." box="[234,500,473,495]" pageId="5" pageNumber="282" pagination="33 - 38" refId="ref18990" refString="MASUMOTO, K., ANDO, Κ. &amp; AKITA, Κ. (2006): New or little known tenebrionid species from Japan (Part 5). A new species belonging to a new genus (Coleoptera: Tenebrionidae: Helopini). - Entomological Review of Japan 61: 33 - 38." type="journal article" year="2006">
Masumoto 
<emphasis id="B966C322FFFFFFF0FEAFA0B757DA39EE" box="[355,395,474,495]" italics="true" pageId="5" pageNumber="282">et al</emphasis>
., 2006: 33
</bibRefCitation>
, figs 1–9; 
<bibRefCitation id="EF8362C1FFFFFFF0FD96A0B755F139EE" author="NABOZHENKO, M. V. &amp; LOBL, I." box="[602,928,473,496]" pageId="5" pageNumber="282" pagination="241 - 257" refId="ref19945" refString="NABOZHENKO, M. V. &amp; LOBL, I. (2008) Tribe Helopini. Pp. 241 - 257. In: LOBL, I. &amp; SMETANA, A. (eds): Catalogue of Palearctic Coleoptera. Vol. 5. Tenebrionoidea. Apollo Books, Stenstrup." type="book chapter" year="2008">Nabozhenko &amp; Löbl, 2008: 253</bibRefCitation>
.
</paragraph>
<paragraph id="8BAD1F30FFFFFFF0FF26A09A54F63A63" blockId="5.[173,1127,473,697]" pageId="5" pageNumber="282">
Type material examined. 
<materialsCitation id="3B7A156DFFFFFFF0FDC8A09B54F63A63" collectingDate="2004-05-28" collectionCode="NMNS, T" country="Japan" location="Mt. Sangai-yama" pageId="5" pageNumber="282" specimenCode="R4" specimenCount="2" stateProvince="Ishikawa" typeStatus="holotype">
<typeStatus id="54A9A192FFFFFFF0FDC8A09B543A3A0D" box="[516,619,502,524]" pageId="5" pageNumber="282" type="holotype">Holotype</typeStatus>
: m (
<collectionCode id="ED0387F5FFFFFFF0FD56A09A54B63A0C" box="[666,743,503,525]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="5" pageNumber="282">NMNS</collectionCode>
) (separated specimen, placed on 5 plates) with labels: “ 
<typeStatus id="54A9A192FFFFFFF0FE4AA37E57BD3A28" box="[390,492,531,553]" pageId="5" pageNumber="282" type="holotype">Holotype</typeStatus>
<emphasis id="B966C322FFFFFFF0FE3FA37E54B53A29" box="[499,740,531,552]" italics="true" pageId="5" pageNumber="282">
<taxonomicName id="4C1264B3FFFFFFF0FE3FA37E54D23A29" box="[499,643,531,552]" class="Insecta" family="Tenebrionidae" genus="Nipponohelops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="282" phylum="Arthropoda" rank="genus">Nipponohelops</taxonomicName>
ishikawai
</emphasis>
Ms., And. &amp; Aki.”, “ 
<collectingCountry id="F3055FA0FFFFFFF0FC0CA37E52513A29" box="[960,1024,531,552]" name="Japan" pageId="5" pageNumber="282">Japan</collectingCountry>
, Honshu, Nagano-ken, Takatô-machi, 
<location id="8ECD49EBFFFFFFF0FE16A35D54C13A47" LSID="urn:lsid:plazi:treatment:03BBAE26FFFFFFF0FEADA1D3548D3AB8:8ECD49EBFFFFFFF0FE16A35D54C13A47" box="[474,656,560,582]" country="Japan" name="Mt. Sangai-yama" pageId="5" pageNumber="282" stateProvince="Ishikawa">Mt. Sangai-yama</location>
, 
<date id="FFAC39F0FFFFFFF0FD57A35D55503A44" box="[667,769,560,581]" pageId="5" pageNumber="282" value="2004-05-28">
<collectingDate id="EFE8C018FFFFFFF0FD57A35D55503A44" box="[667,769,560,581]" pageId="5" pageNumber="282" value="2004-05-28">28.V.2004</collectingDate>
</date>
, H. 
<collectingRegion id="49D6D1D2FFFFFFF0FCE0A35D55DD3A47" box="[812,908,560,582]" country="Japan" name="Ishikawa" pageId="5" pageNumber="282">Ishikawa</collectingRegion>
leg.”, “ 
<date id="FFAC39F0FFFFFFF0FC15A35D52193A47" box="[985,1096,560,582]" pageId="5" pageNumber="282" value="2004-05-28">
<collectingDate id="EFE8C018FFFFFFF0FC15A35D52193A47" box="[985,1096,560,582]" pageId="5" pageNumber="282" value="2004-05-28">2004.05.28</collectingDate>
</date>
上 OiDZDzdzẘoi H. 
<collectingRegion id="49D6D1D2FFFFFFF0FEA8A32057953A62" box="[356,452,589,611]" country="Japan" name="Ishikawa" pageId="5" pageNumber="282">Ishikawa</collectingRegion>
leg. 
<specimenCode id="DBB4B74BFFFFFFF0FE3AA32054443A63" box="[502,533,589,610]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="282">R4</specimenCode>
. 
<collectionCode id="ED0387F5FFFFFFF0FDD2A32054653A60" box="[542,564,589,609]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="282">T</collectionCode>
ḩvẉẅ”.
</materialsCitation>
</paragraph>
<paragraph id="8BAD1F30FFFFFFF0FF26A30757D23A9D" blockId="5.[173,1127,473,697]" pageId="5" pageNumber="282">
<materialsCitation id="3B7A156DFFFFFFF0FF26A307524C3A7E" box="[234,1053,617,640]" collectionCode="CKA" county="Mt. Mitsugai-san" elevation="1" location="Ina City" municipality="Takato" pageId="5" pageNumber="282" specimenCount="1" stateProvince="Nagano">
Material examined. 
<quantity id="4CEAB2D5FFFFFFF0FE70A307578F3A7E" box="[444,478,617,639]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="282" unit="m" value="1.0">
<elevation id="003FF803FFFFFFF0FE70A307578F3A7E" box="[444,478,617,639]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="282" unit="m" value="1.0">1 m</elevation>
</quantity>
(
<collectionCode id="ED0387F5FFFFFFF0FE27A30754723A81" box="[491,547,618,640]" pageId="5" pageNumber="282">CKA</collectionCode>
), 
<collectingCounty id="62CC67BCFFFFFFF0FDFEA30754B83A81" box="[562,745,618,640]" pageId="5" pageNumber="282">Mt. Mitsugai-san</collectingCounty>
, 
<collectingMunicipality id="6BC9854AFFFFFFF0FD3EA307556A3A81" box="[754,827,618,640]" pageId="5" pageNumber="282">Takatô</collectingMunicipality>
, 
<location id="8ECD49EBFFFFFFF0FC88A30755CB3A81" LSID="urn:lsid:plazi:treatment:03BBAE26FFFFFFF0FEADA1D3548D3AB8:8ECD49EBFFFFFFF0FC88A30755CB3A81" box="[836,922,618,640]" county="Mt. Mitsugai-san" municipality="Takato" name="Ina City" pageId="5" pageNumber="282" stateProvince="Nagano">Ina City</location>
, 
<collectingRegion id="49D6D1D2FFFFFFF0FC6FA30755A83A7E" box="[931,1017,618,639]" country="Japan" name="Nagano" pageId="5" pageNumber="282">Nagano</collectingRegion>
, 
<date id="FFAC39F0FFFFFFF0FBCEA304524C3A7E" box="[1026,1053,617,639]" pageId="5" pageNumber="282" value="2010-05-15">15.</date>
</materialsCitation>
<materialsCitation id="3B7A156DFFFFFFF0FBD0A304572E3A9D" collectionCode="V" collectorName="Kiyoshi Ando" pageId="5" pageNumber="282" specimenCount="1" stateProvince="Nagano">
<collectionCode id="ED0387F5FFFFFFF0FBD0A304527E3A7E" box="[1052,1071,617,639]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="282" type="Museum">V</collectionCode>
.2010 (leg. 
<collectorName id="26E77AE6FFFFFFF0FF2DA3EB57293A9D" box="[225,376,646,668]" pageId="5" pageNumber="282">Kiyoshi Ando</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD1F30FFFFFFF0FF26A3CE548D3AB8" blockId="5.[173,1127,473,697]" box="[234,732,675,697]" pageId="5" pageNumber="282">
Distribution. 
<collectingCountry id="F3055FA0FFFFFFF0FEB4A3C957E63AB8" box="[376,439,676,697]" name="Japan" pageId="5" pageNumber="282">Japan</collectingCountry>
: Honshu (Central districts).
</paragraph>
</subSubSection>
</treatment>
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