A revision of the Chilean Brachyglutini - Part 4. Revision of Achilia Reitter, 1890: A. puncticeps and A. approximans species groups, with description of seven new species (Coleoptera: Staphylinidae: Pselaphinae) Author Sabella, Giorgio Author Cuccodoro, Giulio Author Kurbatov, Sergey A. text Revue suisse de Zoologie 2019 2019-03-31 126 1 127 145 journal article 10.5281/zenodo.2619528 a0525610-166a-4a90-97cb-6cd78d98a706 0035-418 2619528 Achilia reitteri n. sp. Figs 2 , 9 , 14 , 21 , 31-32 , 57 Holotype : MHNG (# MHNG-ENTO-1383); 1 ♂; SOUTHERN CHILI : Región Los Lagos : Osorno prov. : Puyehue National Park , Antillanca road; 500-1000 m ; 18-20.XII.1984 ; S. & J. Peck ; car netting. Paratypes (96): SOUTHERN AND CENTRAL CHILI : Región Aysén : Aysén prov. : MHNG ; 1 ♂; 16 km NW Cisnes Medio , Río Grande ; 200 m ; 30.XII.1984 - 28.I.1985 ; S. & J. Peck; FIT mature beech forest . – MHNG ; 1 ♂; Rio Simpson National Park , 33 km E Puerto Aysén ; 70 m ; 31.XII.1984 / 26.I.1985 ; S. & J. Peck; FIT, select cut forest. – Región Los Lagos : Llanquihue prov.: UNHC ; 1 ♂; Salto Petrohué , 6.4 km SW Petrohué ; 140 m ; 28.XII.1982 ; A. Newton & M. Thayer ; Valdivian rainforest, Berlese , leaf & log litter, forest floor . – FMNH ; 1 ♂; Lago Chapo , 13.5 km E Correntoso , site 656; 310 m ; 16-27.XII.1982 ; A. Newton & M. Thayer; Valdivian rainforest, flight intercept (windows) trap . – FMNH ; 1 ♀ ; same data, but Berlese , leaf & log litter, forest floor . – Chiloé prov.: FMNH ; 1 ♂; Ahoni Alto ; 70 m ; 22.II.1988 ; L. Masner, primary forest . – Osorno prov.: UNHC ; 1 ♂; 7.7 km NE Termas de Puyehue , site 664; 200 m ; 19-25.XII.1982 ; A. Newton & M. Thayer ; Valdivian rainforest, Berlese , leaf & log litter, forest floor . – FMNH ( FMHD #85- 923 , # 85-38 ); 3 ♂; Puyehue National Park , Antillanca road; 500 -1000 m; 18-20.XII.1984 ; S. & J. Peck; car netting . – MHNG ; 24 ♂ and 38 ♀ ; same data . – MHNS, 1 ♂ and 1 ♀ ; same data . – FMNH ; 3 ♂ and 3 ♀ ; Puyehue National Park , Antillanca road, site 659; 720 m ; 18-24.XII.1982 ; A. Newton & M. Thayer; Nothofagus spp. forest, flight intercept . – UNHC ; 2 ♂ and 2 ♀ ; same data . – FMNH ; 1 ♀ ; Puyehue National Park , Antillanca road; 470 -720 m; 18-24.XII.1982 ; A. Newton & M. Thayer ; Valdivian rainforest, screen sweeping at dusk . – MHNG ; 5 ♂; Puyehue National Park , Antillanca road; 600 -1000 m; 20.XII.1984 ; S. & J. Peck; car netting . – MHNG ; 1 ♂; Puyehue National Park , Aguas Calientes ; 500 m ; 20.XII.1984 ; S. & J. Peck; forest litter on trail, sifting . – MHNG ; 1 ♂; Puyehue National Park , Aguas Calientes , station 25a; 400 -500 m; 31.XII.1990 / 01.I.1991 ; D. Agosti & D. Burckhardt . – FMNH ( FMHD # 97-5 ); 1 ♂; Puyehue National Park , 4 km E Anticura ; 40° 39.73’S 72° 08.10’W ; 460 m ; 30.I.1997 ; A. Newton & M. Thayer 985-3; Valdivian rainforest w/large, Saxegothea , flight intercept trap. – Región Los Ríos : Valdivia prov.: MHNG ; 1 ♂; Alerce Costiero National Park , near Chaihuin ; 500 m ; 15.II.2018 ; G. Sabella & D. Mifsud; forest litter . – Región Araucanía : Malleco prov.: MHNG ; 1 ♂; Puren , Contulmo Natural Monument ; 350 m ; 11.XII.1984 - 13.II.1985 ; S. & J. Peck 85-16; FIT, mixed evergreen forest & MT Nothofagus . – Región Bío Bío: Concepción prov.: MSNG ; 1 ♂; El Manzano ; TC-189; 03.I.1988 ; T. Cekalovic . Figs 7-10. Aedeagi (7-8) and mesotibiae (9-10) of Achilia species. (7) A . cunniceps n. sp. (8, 10) A. zaurda n. sp. (9) A. reitteri n. sp. Figs 11-14. Antennae of Achilia species. (11) A . cunniceps n. sp. (12) A. baburra n. sp. (13) A. puncticeps . (14) A. reitteri n. sp. Description: Body 1.30-1.40 mm long, entirely dark brown with reddish elytra darker at base and along sutural stria; antennae, palpi, and legs reddish-brown. Head with eyes moderately protruding, distinctly shorter than slightly convex temples. Pronotum as wide as head; median antebasal fovea as large as lateral ones. First abdominal tergite with basal striae extending to about one-quarter of paratergal length, and separated at base by more than one-quarter of tergal width. Male : Head as in Figs 31-32 ; frons flattened, densely punctate and pubescent, disc with broad V-shaped depression with distinctly raised margins; vertexal sulcus shallowly impressed; frontal lobe wide and flattened. Antennae ( Fig. 14 ) with scape and pedicel longer than wide; antennomere III as long as wide, antennomere IV wider than long; antennomere V longer than wide, antennomeres VI-VIII wider than long, VII bearing two long setae on its mesal margin; antennomere IX transverse with protruding distal mesal margin and bearing setae on its mesal margin; antennomere X strongly transverse: antennomere XI very elongate and distinctly longer than VI-X combined, bearing long and thin subbasal seta inserted in deep impression, its surface with many tubercles. Metaventrite raised at middle, this area with some fine punctures, distally pubescent, and divided by wide median sulcus. Legs with trochanters very elongate; surface of protrochanters and mesotrochanters bearing numerous long bristles; ventral margin of mesotrochanters projecting posteriorly as spine ( Fig. 21 ); profemora and mesofemora slightly thickened; mesotibiae with short subapical spur on medial margin ( Fig. 9 ), distal half slightly sinuate bearing long and thick bristles and long and wide macroseta; distal half of metatibiae slightly sinuate. Abdominal tergites and ventrites unmodified. Aedeagus ( Fig. 2 , dorsal longitudinal struts not shown) 0.27-0.30 mm long; long bifid medial sclerites associated on each side with four pointed sclerites. Figs 15-18. Antennae of Achilia species. (15) A . nipponobythoides n. sp. (16) A. adorabilis n. sp. (17) A. trulla n. sp. (18) A. zaurda n. sp. Female : Similar to male except: head with frons slightly convex (not flattened) without punctures and with large vertexal fovea beside each eye; vertexal sulcus present, well-impressed; antennae shorter and thinner than male; metaventrite, and legs unmodified. Figs 19-28. Protrochanters (19-20) and mesotrochanters (21-28) of Achilia species. (19, 22) A. baburra n. sp. (20, 28) A . nipponobythoides n. sp. (21) A. reitteri n. sp. (23) A . cunniceps n. sp. (24) A. trulla n. sp. (25) A. puncticeps . (26) A. zaurda n. sp. (27) A . adorabilis n. sp. Figs 29-34. (29-30) Achilia puncticeps . (31-32) A. reitteri n. sp. (33-34) A. baburra n. sp. Male head in (29, 31, 33) dorsal and (30, 32, 34) lateral views. Figs 35-40. (35, 37, 39) Achilia cunniceps n. sp. (36, 38, 40) A. nipponobythoides n. sp. Male head in (35-36) dorsal, (37-38) lateral and (39-40) frontal views. Collecting data: Collected from December to February, mainly in Valdivian rainforests, at elevations ranging from 70 m to 1000 m . Most specimens came from car netting and flight intercept traps, but also by sifted samples of leaf and log litter. Distribution: Achilia reitteri n. sp. is known from Southern and Central Chile ( Fig. 57 : green diamonds), ranging from Aysén to Concepción provinces. Comments: Achilia reitteri n. sp. is similar to A. puncticeps from which it is easily distinguished by the male features of the head (compare Figs 29-30 and 31-32 ), antennae (compare Figs 13 and 14 ), mesotrochanters (compare Figs 21 and 25 ), and the copulatory pieces of the aedeagus (compare Figs 1 and 2 ). The females of the two species are distinguished because A. puncticeps has the sides of the frontal lobe clearly narrowed (slightly narrowed in A. reitteri n. sp. )