The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida) Author Golovatch, Sergei Author Evsyukov, Aleksandr Author Reip, Hans text Zootaxa 2016 2016-03-01 4085 1 1 51 journal article 31448 10.11646/zootaxa.4085.1.1 429cf011-ec4c-4a61-b894-2bd32b2ffc9b 1175-5326 1052381 7F819439-FD77-4EF2-8E26-7B46F9B5FAAB Brachydesmus ( Haplobrachidesmus ) simplex sp. n. Fig. 29 , Map 3 Material examined. Holotype male (ZMUM p3037), ABKHAZIA, Gagra Distr., Lake Ritsa, 950 1100 m a.s.l., Fagus , Abies , Picea , Acer etc. forest, litter, under bark and stones, 13 14.VIII.1986 , leg. S. Golovatch. Paratypes: 5 males (ZMUM p3038), same data, together with holotype; 1 male, 6 females (ZMUM p3039), Myussera Nature Reserve, 20 130 m a.s.l., Castanea , Alnus etc. forest, 8 10.IV.1983 , leg. S. Golovatch; 3 males (MIZW), 1 male (ZMUM p3040), RUSSIA, Krasnodar Prov., Sochi, open Carpinus and Quercus forest, loam, 7.XII.1963 ; 1 male (MIZW), same locality, dendrarium and city park, under stones, 9.XII.1963 , all leg. P. Pisarski. Diagnosis. See Diagnosis of B. kvavadzei sp. n. , as well as Key below. Name. To emphasize the particularly simple gonopod exomere. Description. Length 6 7.5 mm , width of midbody metazonae 0.8 1.0 mm (male, female), metazonite to prozonite width ratio ca 1.5 1.6 ( Fig. 29A ). Coloration uniformly pallid to light yellow. All characters as in B. assimilis ( Fig. 29A ), except as follows. In width, collum <2 <3 <head = 4 <5. Paraterga well-developed, their fore margins nearly straight, while anterolateral corners clearly angular; caudolateral corner mostly acute-angled and dentiform starting with segment 5, subspiniform and drawn behind rear tergal margin starting with segment 15 ( Fig. 29A ). Tergal setae mostly very short and subclavate. Metatergal sculpture indistinct, but at least middle and caudal rows of flat small tubercles visible. Male legs clearly incrassate compared to female, but male prefemora not bulging laterad; sphaerotrichomes absent ( Fig. 29B ). Gonopod ( Fig. 29C 29G ) bipartite, stout, exomere with only a single, small, apical tooth ( e ).