<document id="D3F7AE85259F4A18FF8219EEF8E064C3" ID="10.11646/zootaxa.4138.3.2" ID-DOI="10.11646/zootaxa.4138.3.2" ID-GBIF-Dataset="c785c9ac-b71c-4487-b770-35ebfe797c8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="258464" ID-ZooBank="DDF68118-EA96-49A5-B316-47814E19ED3D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468823189128" checkinUser="plazi" docAuthor="Leschen, Richard A. B., Escalona, Hermes E. &amp; Elgueta, Mario" docDate="2016" docId="746C8797FFEAD015ACDDFC61FB1192DE" docLanguage="en" docName="zootaxa.4138.3.2.pdf" docOrigin="Zootaxa 4138 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trachyderas Philippi &amp; Philippi 1864" docType="treatment" docVersion="9" lastPageNumber="469" masterDocId="8855FFEFFFF6D009AC4AFF96FF8B9636" masterDocTitle="Phylogeny of the Gondwanan beetle family Ulodidae (Tenebrionoidea)" masterLastPageNumber="473" masterPageNumber="441" pageNumber="469" updateTime="1698365200336" updateUser="plazi">
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<mods:title id="F782CD496DE4821055FC390B14371E23">Phylogeny of the Gondwanan beetle family Ulodidae (Tenebrionoidea)</mods:title>
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<mods:namePart id="82C670DBB4D2FDCA693F230D49E5325B">Leschen, Richard A. B.</mods:namePart>
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<mods:namePart id="1FBBE00A60121F691E36FC8FB9B821F1">Escalona, Hermes E.</mods:namePart>
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<mods:namePart id="0B5C954C7E426AC29B6BBA1B2939E69E">Elgueta, Mario</mods:namePart>
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<mods:date id="EE52620C3C18951C0D9B5ABD481C1255">2016</mods:date>
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<taxonomicName id="3BC54D02FFEAD015ACDDFC61FD1A9226" authority="Philippi &amp; Philippi, 1864: 349" authorityName="Philippi &amp; Philippi" authorityPageNumber="349" authorityYear="1864" box="[151,657,1015,1040]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="CEB1EA93FFEAD015ACDDFC61FEAC9226" bold="true" box="[151,295,1015,1040]" italics="true" pageId="28" pageNumber="469">Trachyderas</emphasis>
<treatmentCitation id="7D641090FFEAD015AD67FC61FD1A9226" author="Philippi" box="[301,657,1015,1040]" page="349" pageId="28" pageNumber="469" year="1864">
<bibRefCitation id="98544B70FFEAD015AD67FC61FDD09226" author="Philippi" box="[301,603,1015,1040]" pageId="28" pageNumber="469" refString="Philippi, R. A. &amp; Philippi, F. H. E. (1864) Beschreibung einiger neuen Chilenischen Kafer. Stettiner Entomologische Zeitung, 25 (10 - 12), 313 - 406." type="journal article" year="1864">Philippi &amp; Philippi, 1864</bibRefCitation>
: 349
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–350
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</paragraph>
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<paragraph id="FC7A3681FFEAD015ACDDFB8BFCEB926E" blockId="28.[151,1436,1015,1256]" pageId="28" pageNumber="469">
<typeStatus id="237E8823FFEAD015ACDDFB8BFF449203" box="[151,207,1053,1077]" pageId="28" pageNumber="469">Type</typeStatus>
species: 
<taxonomicName id="3BC54D02FFEAD015AD7EFB8AFC229203" authority="Philippi &amp; Philippi, 1864: 350" authorityName="Philippi &amp; Philippi" authorityPageNumber="350" authorityYear="1864" box="[308,937,1052,1077]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="CEB1EA93FFEAD015AD7EFB8AFDDB9203" box="[308,592,1052,1077]" italics="true" pageId="28" pageNumber="469">Trachyderas cancellatum</emphasis>
<bibRefCitation id="98544B70FFEAD015AE1DFB8AFCFE9203" author="Philippi" box="[599,885,1052,1077]" pageId="28" pageNumber="469" refString="Philippi, R. A. &amp; Philippi, F. H. E. (1864) Beschreibung einiger neuen Chilenischen Kafer. Stettiner Entomologische Zeitung, 25 (10 - 12), 313 - 406." type="journal article" year="1864">Philippi &amp; Philippi, 1864</bibRefCitation>
: 350
</taxonomicName>
(junior synonym of 
<taxonomicName id="3BC54D02FFEAD015A8C4FB8AFE74926E" authority="Fairmaire &amp; Germain, 1860: 2" authorityName="Fairmaire &amp; Germain" authorityPageNumber="2" authorityYear="1860" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="species" species="costulatum">
<emphasis id="CEB1EA93FFEAD015A8C4FB8AFA179203" box="[1166,1436,1052,1077]" italics="true" pageId="28" pageNumber="469">Trachyderas costulatum</emphasis>
(
<bibRefCitation id="98544B70FFEAD015ACD4FBA9FE7D926E" author="Fairmaire" box="[158,502,1087,1112]" pageId="28" pageNumber="469" refString="Fairmaire, L. &amp; Germain, P. (1860) Coleoptera Chilensia. Typographie F. Malteste et cie. Part 2, Section 1, 8 pp." type="book" year="1860">Fairmaire &amp; Germain, 1860: 2</bibRefCitation>
)
</taxonomicName>
[
<taxonomicName id="3BC54D02FFEAD015AE45FBA9FD2F926E" box="[527,676,1087,1112]" class="Insecta" family="Tenebrionidae" genus="Bolitophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="CEB1EA93FFEAD015AE45FBA9FD2F926E" box="[527,676,1087,1112]" italics="true" pageId="28" pageNumber="469">Bolitophagus</emphasis>
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]), by monotypy.
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<paragraph id="FC7A3681FFEAD015ACDDFBF2FEFA924B" blockId="28.[151,1436,1015,1256]" box="[151,369,1124,1149]" pageId="28" pageNumber="469">
Distribution: 
<collectingCountry id="84D27611FFEAD015AD67FBF2FEE6924B" box="[301,365,1124,1149]" name="Chile" pageId="28" pageNumber="469">Chile</collectingCountry>
.
</paragraph>
<paragraph id="FC7A3681FFEAD015ACDDFB11FB1192DE" blockId="28.[151,1436,1015,1256]" pageId="28" pageNumber="469">
Included species (1). 
<taxonomicName id="3BC54D02FFEAD015ADCDFB1FFD959296" box="[391,542,1159,1184]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="species" species="costulatum">
<emphasis id="CEB1EA93FFEAD015ADCDFB1FFD959296" box="[391,542,1159,1184]" italics="true" pageId="28" pageNumber="469">T. costulatum</emphasis>
</taxonomicName>
(
<bibRefCitation id="98544B70FFEAD015AE66FB11FCE09296" author="Fairmaire" box="[556,875,1159,1184]" pageId="28" pageNumber="469" refString="Fairmaire, L. &amp; Germain, P. (1860) Coleoptera Chilensia. Typographie F. Malteste et cie. Part 2, Section 1, 8 pp." type="book" year="1860">Fairmaire &amp; Germain, 1860</bibRefCitation>
: 2 [
<taxonomicName id="3BC54D02FFEAD015AFDFFB11FBF79296" authority="Illiger" authorityName="Illiger" box="[917,1148,1159,1184]" class="Insecta" family="Tenebrionidae" genus="Bolitophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="CEB1EA93FFEAD015AFDFFB11FBA19296" box="[917,1066,1159,1184]" italics="true" pageId="28" pageNumber="469">Bolitophagus</emphasis>
Illiger
</taxonomicName>
], combination by 
<bibRefCitation id="98544B70FFEAD015A90EFB11FF5892F2" author="Gebien" pageId="28" pageNumber="469" refString="Gebien, H. (1939) Katalog der Tenebrioniden. Teil II (part). Mitteilungen der Munchener Entomologischen Gesellschaft, 29, 443 - 474 (466 - 497), 739 - 770 (498 - 529)." type="journal article" year="1939">Gebien, 1939</bibRefCitation>
: 759 as 
<taxonomicName id="3BC54D02FFEAD015AD67FB3AFDBA92F3" box="[301,561,1196,1221]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="CEB1EA93FFEAD015AD67FB3AFDBA92F3" box="[301,561,1196,1221]" italics="true" pageId="28" pageNumber="469">Trachyderas costulatus</emphasis>
</taxonomicName>
and noted with an “a.” = aberration of 
<taxonomicName id="3BC54D02FFEAD015AFA3FB3AFBF992F3" box="[1001,1138,1196,1221]" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="CEB1EA93FFEAD015AFA3FB3AFBF992F3" box="[1001,1138,1196,1221]" italics="true" pageId="28" pageNumber="469">cancellatum</emphasis>
</taxonomicName>
; = 
<taxonomicName id="3BC54D02FFEAD015A8D2FB3BFE0592DE" authority="Philippi &amp; Philippi, 1864: 350" authorityName="Philippi &amp; Philippi" authorityPageNumber="350" authorityYear="1864" class="Insecta" family="Ulodidae" genus="Trachyderas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="469" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="CEB1EA93FFEAD015A8D2FB3BFAB792F3" box="[1176,1340,1196,1221]" italics="true" pageId="28" pageNumber="469">T. cancellatum</emphasis>
<bibRefCitation id="98544B70FFEAD015A909FB3AFEDD92DE" author="Philippi" pageId="28" pageNumber="469" refString="Philippi, R. A. &amp; Philippi, F. H. E. (1864) Beschreibung einiger neuen Chilenischen Kafer. Stettiner Entomologische Zeitung, 25 (10 - 12), 313 - 406." type="journal article" year="1864">Philippi &amp; Philippi, 1864</bibRefCitation>
: 350
</taxonomicName>
, combination by 
<bibRefCitation id="98544B70FFEAD015AE05FB59FCAE92DE" author="Blackwelder" box="[591,805,1231,1256]" pageId="28" pageNumber="469" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185 (part 3), 343 - 550." type="journal article" year="1945">Blackwelder, 1945</bibRefCitation>
: 526, see also Peña, 1966a: 437).
</paragraph>
</subSubSection>
</treatment>
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