Review of the Prabhasa Moore, 1878 genus-group (Lepidoptera, Erebidae, Arctiinae)
Author
Dubatolov, Vladimir V.
Author
Volynkin, Anton V.
Author
Kishida, Yasunori
text
Zootaxa
2018
2018-04-11
4407
3
383
400
journal article
30294
10.11646/zootaxa.4407.3.6
d6a4aaf4-4045-4302-8220-413a7eca4f99
1175-5326
1216616
3FF82287-8A03-4FE6-803D-EC8C2A370B6D
Macohasa
Dubatolov, Volynkin & Kishida
,
gen. nov.
Type species:
Macohasa cernyi
Dubatolov, Volynkin & Kishida
,
sp. nov.
Macotasa:
HolloWay 2001
: 305
.
Prabhasa
: Černý & Pinratana 2009: 139
.
Description.
Forewing shape resembles that of
Eilema
, but dorsum noticeably conVeX in its basal half; its outer edge folded inwards, stronger in males. Costal Vein with androconial scales in basal half.
Male genitalia
: uncus long, narrow, with terminal denticle; scaphium heaVily sclerotized; apeX of ValVa broad, without distal process, costal margin straight; medial area of ValVe with a harpe-like fold; sacculus short, with bented apeX; aedeagus short, Vesica with seVeral diVerticula.
Female genitalia
: oVipositor short and broad; apophyses long and thin; ductus bursae, narrow, short, membranous, with a large, heaVily sclerotized plate Ventrally; corpus bursae large, membranous, with one signum; appendiX bursae short, bulbous, situated latero-posteriorly.
FIGURES 1–10.
Prabhasa
and
Macohasa
spp.: adults. 1,
P. venosa
, ♂ syntype, Darjeeling (©NHMUK); 2,
P. venosa
, ♀ syntype, Darjeeling (©ZFMK); 3,
P. venosa
, ♂, [India, Meghalaya] Khasis (©NHMUK); 4,
M. cernyi
, holotype ♂, N Vietnam, Bac Giang prov. (SZMN); 5,
M. cernyi
, paratype ♂, Thailand, Kanchanaburi prov. (CKC); 6,
M. cernyi
, paratype ♀, N Vietnam, Dien Bien prov. (MWM/ZSM); 7,
M. cernyi
, paratype ♂, Thailand, Phetchabun prov. (CKC); 8,
M. cernyi
, paratype ♀, Thailand, Phetchabun prov. (CKC); 9,
M. orientalis
, holotype ♂, Borneo (©NHMUK); 10,
M. orientalis
, holotype ♀ of
pentaspila
, Singapore (©NHMUK).
FIGURES 11–19.
Macohasa
and
Chinasa
spp.: adults. 11,
M. orientalis
, ♂, Malaysia, Malacca Peninsula (MWM/ZSM); 12,
M. orientalis
, ♀, Malaysia, Malacca Peninsula (MWM/ZSM); 13,
M. tetraspila
, holotype ♂, Thailand, Chantaburi prov. (MWM/ZSM); 14,
M. tetraspila
, paratype ♀, Thailand, Chantaburi prov. (MWM/ZSM); 15,
M. dimorpha
, syntype ♂, Philippines, Luzon (©NHMUK); 16,
M. dimorpha
, syntype ♀, Philippines, Luzon (©NHMUK); 17,
Ch. costalis costalis
, holotype ♂, N China (©NHMUK); 18,
Ch. costalis costalis
, ♂, China, Zhejiang (MWM/ZSM); 19,
Ch. costalis costalis
, ♀, China, Guangdong (SZMN).
FIGURES 20–23.
Chinasa
spp.: adults. 20,
Ch. costalis bicoloriceps
, ♂, Taiwan, Kaohsiung prov. (MWM/ZSM); 21,
Ch. costalis bicoloriceps
, ♀, Taiwan, Nan-Tou prov. (MWM/ZSM); 22,
Ch. sapa
, holotype ♂, N Vietnam (MWM/ZSM); 23,
Ch. sapa
, paratype ♀, N Vietnam (MWM/ZSM).
Remarks. 1.
The new genus differs significantly either from species of
Prabhasa
, or
Zadadra
(
type
species
Zadadra distorta
(
Moore, 1872
)
(
Fig. 30
)); broad and weakly sclerotized apeX of cucullus is not characteristic for both genera (only “
Zadadra
”
cucullata
Joshi, Kirti & Singh, 2015
has broad distal part of ValVe, but it is heaVily sclerotized (see:
Joshi
et al.
2015
: figs. 2–4), and the generic placement of this species will be clarified in the further reVision of the genus
Zadadra
); harpe-like fold is also an autapomorphic character of the genus.
2.
Macohasa orientalis
(
Hampson, 1905
)
,
comb. nov.
,
Macohasa tetraspila
(Černý, 2009)
,
comb. nov.
and
Macohasa dimorpha
(
Hampson, 1918
)
,
comb. nov.
were included formerly in
Macotasa
(
Holloway 1982
;
2001
;
Fang 2000
; Černý & Pinratana 2009; Bucsek 2012), but they are clearly not congeneric to the
type
species of
Macotasa
,
M. biplagella
(Butler, 1877)
(
Figs. 31
,
41
): in the male genitalia,
M. biplagella
has the bifid uncus, the furcated juXta, the large basal saccular process, and the small sclerotized harpe in the medial section of ValVe, whereas species of
Macohasa
haVe a simple uncus, a plate-like juXta, no basal saccular process, and no harpe (but a membranous fold in the medial section of ValVe is presented); in the female genitalia,
M. biplagella
has no sclerotized Ventral plate in ductus bursae which is characteristic for species of
Macohasa
. Another character of
Macotasa
, “a fold in the cell on male forewing ridges the dorsal surface, the basal part of this ridge coVered by an area of coarse scales to just beyond the ridge” (
Holloway 1982
;
2001
), is not autapomorphic, but common in
Macohasa
,
Prabhasa, Chinasa
,
some
Teulisna
Walker, 1862
, genera with quite different male genitalia.
Etymology.
The genus name is a combination of the generic names
Macotasa
and
Prabhasa
. Gender is feminine.