Redescription of three Hippasa species from China (Araneae: Lycosidae), with a proposed species group-division and diagnosis
Author
Wang, Lu-Yu
Author
Li, Zong-Xu
Author
Zhou, Ke-Xin
Author
Zhang, Zhi-Sheng
text
Zootaxa
2015
3974
2
231
244
journal article
10.11646/zootaxa.3974.2.7
17d9c385-2450-44cf-9f3e-ddc970096f57
1175-5326
243939
70C27916-E17A-4E97-B2DD-356775A66D65
Hippasa
partita-
group
Diagnosis.
In the
H. partita
species group, the bifurcation of the two arms of the median apophysis is usually located proximally and may even be invisible in the ventral view of the palp (
Figs 7
A, 8C). The anterior arm (
AA
) is curved and usually sickle or hook-shaped (
Figs 7
B, 8D). The epigyne lacks a scape but has an obvious atrium (
Figs 7
C, 8E–F).
Composition and distribution.
Seventeen species are included (
Fig. 2
).
FIGURE 2.
Distribution records of the
Hippasa partita
-group.
Ten of them found only in Africa:
H. affinis
Lessert, 1933
(♀, from
Angola
)
H. albopunctata
Thorell, 1899
(♀♂, from
Cameroon
and
Ivory Coast
)
H. australis
Lawrence, 1927
(♀♂, from Southern Africa)
H. brechti
Alderweireldt & Jocqué, 2005
(♀, from
Ivory Coast
and
Togo
)
H. cinerea
Simon, 1898
(♀♂, from Africa)
H. decemnotata
Simon, 1910
(♀♂, from West Africa)
H. elienae
Alderweireldt & Jocqué, 2005
(♀, from
Tanzania
and
Kenya
)
H. funerea
Lessert, 1925
(♀♂, from Southern Africa and
Botswana
)
H. innesi
Simon, 1889
(♀, from
Egypt
)
H. marginata
Roewer, 1960
(
Roewer 1960a, ♂, from
Cameroon
)
Two of them from both Africa and Asia:
H. partita
(
O. Pickard-Cambridge, 1876
)
(♀♂, from
Egypt
,
Saudi Arabia
,
Pakistan
,
India
and
Bangladesh
)
H. sinai
Alderweireldt & Jocqué, 2005
(♂, from
Egypt
and
Saudi Arabia
)
Five of them from Asia only:
H. hansae
Gajbe & Gajbe, 1999
(♀, from
India
)
H. himalayensis
Gravely, 1924
(♀, from
India
)
H. lycosina
Pocock, 1900
(♀♂, from
India
,
Laos
and
China
)
H. olivacea
(
Thorell, 1887
)
(♀, from
Myanmar
)
H. valiveruensis
Patel & Reddy, 1993
(♀, from
India
)
From the distribution data of this species group, it appears that most Asian species were collected from countries near the Indian Ocean. We hypothesise that the ancestors of this group might be from Africa, some of them spread into Asia along the coast of
Egypt
, then through central Asia to South and Southeast Asia, where new species evolved.
Among these 30 species and subspecies, there are 10 species known only from females, two species known only from males and one whose sex is unknown (
H. holmerae sundaica
). Inevitably, some species may be synonymised. For example,
H. lycosina
Pocock, 1900
and
H. olivacea
(
Thorell, 1887
)
share the same distribution and have quite similar epigynes, but the latter is known only from females and has not been revised since the original description. They very likely represent the same species. However, the revision on the whole genus is difficult, especially the species from South Asia, because specimens are unavailable.
Remarks on ungrouped
Hippasa
species.
There are seven species that are not included in the two groups discussed in this paper. Two of them,
H. flavicoma
Caporiacco, 1935
(juvenile, from Karakorum) and
H. simoni
(
Thorell, 1887
)
(♀, from
Myanmar
) have no published figures and only the original descriptions. Judging by the original figures, four species,
H. bifasciata
Buchar, 1997
(♀, from
Bhutan
),
H. fabreae
Gajbe & Gajbe, 1999
(♀, from
India
),
H. haryanensis
Arora & Monga, 1994
(♀♂, from
India
) and
H. wigglesworthi
Gajbe & Gajbe, 1999
(♀, from
India
) are not true
Hippasa
species. Further study or revision based on
type
material is needed. For the remaining species,
H. loundesi
Gravely, 1924
(♀, from
India
), there is insufficient information to decide which group it belongs to.