<document id="06A2B0C9FCA53D5C78CC3F00A9536E6F" ID-CLB-Dataset="31353" ID-DOI="10.11646/zootaxa.4353.2.1" ID-GBIF-Dataset="d93b2bd2-75e1-467c-89d3-e2faee3b10a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1065124" ID-ZooBank="D2902BF9-0213-40F3-91A4-EC4D4FDB3B27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511423577451" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A. &amp; Santos, Carmen Van-Dúnem Neto" docDate="2017" docId="03EFF252F3087F6F14BFA426FC81F86A" docLanguage="en" docName="zootaxa.4353.2.1.pdf" docOrigin="Zootaxa 4353 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius Bonelli 1810" docType="treatment" docVersion="8" lastPageNumber="213" masterDocId="FFD68A2AF3047F631428A273FFF3FFBF" masterDocTitle="Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="213" updateTime="1698503882738" updateUser="ExternalLinkService">
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<mods:title id="11CFA7BA738B671BAE24B0DA1063B1E9">Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="8F3D1CD4748B0525B3A834EF97B37D04">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="092AE68F7F1C90F5D6B1ED148CE59348">Capela, Rúben A.</mods:namePart>
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<mods:namePart id="C4FCD24E1E512F0D1BFF15157E40C16F">Santos, Carmen Van-Dúnem Neto</mods:namePart>
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<treatment id="03EFF252F3087F6F14BFA426FC81F86A" ID-DOI="http://doi.org/10.5281/zenodo.6005636" ID-GBIF-Taxon="137541493" ID-Zenodo-Dep="6005636" LSID="urn:lsid:plazi:treatment:03EFF252F3087F6F14BFA426FC81F86A" httpUri="http://treatment.plazi.org/id/03EFF252F3087F6F14BFA426FC81F86A" lastPageNumber="213" pageId="12" pageNumber="213">
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<paragraph id="8BF94344F3087F6F14BFA426FE48F9D0" blockId="12.[151,443,1621,1647]" box="[151,443,1621,1647]" pageId="12" pageNumber="213">
<heading id="D0B1F428F3087F6F14BFA426FE48F9D0" bold="true" box="[151,443,1621,1647]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<taxonomicName id="4C4638C7F3087F6F14BFA426FE48F9D0" ID-CoL="8KVJR" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[151,443,1621,1647]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F14BFA426FE48F9D0" bold="true" box="[151,443,1621,1647]" pageId="12" pageNumber="213">
<emphasis id="B9329F56F3087F6F14BFA426FEE0F9D0" bold="true" box="[151,275,1621,1647]" italics="true" pageId="12" pageNumber="213">Chlaenius</emphasis>
Bonelli, 1810
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF94344F3087F6F14BFA4EFFC81F86A" blockId="12.[151,1437,1692,2005]" pageId="12" pageNumber="213">
A very speciose genus with fifty two species/subspecies known from Angola (
<bibRefCitation id="EFD73EB5F3087F6F107FA4EFFB10F90A" author="Alves" box="[1111,1251,1692,1717]" pageId="12" pageNumber="213" refString="Alves, M. L. G. (1963) Carabideos de Angola e Mocambique. Memorias da Junta de Investigacoes do Ultramar, 2 Serie, 43, 31 - 74." type="journal article" year="1963">Alves 1963</bibRefCitation>
, 
<bibRefCitation id="EFD73EB5F3087F6F10DEA4EFFA6BF90A" author="Ferreira" box="[1270,1432,1692,1717]" pageId="12" pageNumber="213" refString="Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317." type="journal article" year="1965">Ferreira 1965</bibRefCitation>
, 
<bibRefCitation id="EFD73EB5F3087F6F14BFA4CCFE8DF967" author="Kirschenhofer" box="[151,382,1727,1752]" pageId="12" pageNumber="213" refString="Kirschenhofer, E. (2007) Uber neue und wenig bekannte Chlaenius Arten der Untergattung Lissauchenius MacLeay, 1825 der athiopischen Region (Coleoptera: Carabidae). Acta Coleopterologica, 23 (3), 21 - 32." type="journal article" year="2007">Kirschenhofer 2007</bibRefCitation>
, 
<bibRefCitation id="EFD73EB5F3087F6F15A7A4CCFD80F967" author="Anichtchenko" box="[399,627,1727,1752]" pageId="12" pageNumber="213" refString="Anichtchenko, A. V. (2016) Carabidae of the world. Available from: http: // carabid. org (accessed 9 October 2017)" type="journal article" year="2016">Anichtchenko 2016</bibRefCitation>
, Insectoid.info 2017). Within this assemblage there are one species/ subspecies belonging to sg. 
<taxonomicName id="4C4638C7F3087F6F15FBA497FCA8F942" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" box="[467,859,1764,1789]" class="Insecta" family="Carabidae" genus="Ocybatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F15FBA497FDCEF942" box="[467,573,1764,1789]" italics="true" pageId="12" pageNumber="213">Ocybatus</emphasis>
LaFerté-Sénectère, 1851
</taxonomicName>
, four to sg. 
<taxonomicName id="4C4638C7F3087F6F17CBA497FAC8F942" authority="MacLeay, 1825" authorityName="MacLeay" authorityYear="1825" box="[995,1339,1764,1789]" class="Insecta" family="Carabidae" genus="Lissauchenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F17CBA497FB71F942" box="[995,1154,1764,1789]" italics="true" pageId="12" pageNumber="213">Lissauchenius</emphasis>
MacLeay, 1825
</taxonomicName>
, three to sg. 
<taxonomicName id="4C4638C7F3087F6F1497A574FDFDF89F" authority="Kuntzen, 1919" authorityName="Kuntzen" authorityYear="1919" box="[191,526,1799,1824]" class="Insecta" family="Carabidae" genus="Pachydinodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F1497A574FEAFF89F" box="[191,348,1799,1824]" italics="true" pageId="12" pageNumber="213">Pachydinodes</emphasis>
Kuntzen, 1919
</taxonomicName>
, one to sg. 
<taxonomicName id="4C4638C7F3087F6F16B9A57BFC3BF89F" authority="Waldheim, 1829" authorityName="Waldheim" authorityYear="1829" box="[657,968,1799,1824]" class="Insecta" family="Carabidae" genus="Agostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F16B9A57BFCF7F8A0" box="[657,772,1800,1823]" italics="true" pageId="12" pageNumber="213">Agostenus</emphasis>
Waldheim, 1829
</taxonomicName>
, two to sg. 
<emphasis id="B9329F56F3087F6F1066A57AFB51F89F" box="[1102,1186,1801,1824]" italics="true" pageId="12" pageNumber="213">Epomis</emphasis>
Bonelli, 1810, two to sg. 
<taxonomicName id="4C4638C7F3087F6F1496A55FFDD5F8FB" authority="Boheman, 1848" authorityName="Boheman" authorityYear="1848" box="[190,550,1836,1861]" class="Insecta" family="Carabidae" genus="Rhysotrachelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F1496A55FFE98F8FA" box="[190,363,1836,1861]" italics="true" pageId="12" pageNumber="213">Rhysotrachelus</emphasis>
Boheman, 1848
</taxonomicName>
, two to sg. 
<taxonomicName id="4C4638C7F3087F6F168EA55FFBB0F8FA" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" box="[678,1091,1836,1861]" class="Insecta" family="Carabidae" genus="Tomochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F168EA55FFCD5F8FA" box="[678,806,1836,1861]" italics="true" pageId="12" pageNumber="213">Tomochilus</emphasis>
LaFerté-Sénectère, 1851
</taxonomicName>
, one to sg. 
<taxonomicName id="4C4638C7F3087F6F10EAA55FFEC0F8D7" authority="Jeannel, 1949" authorityName="Jeannel" authorityYear="1949" class="Insecta" family="Carabidae" genus="Pseudochlaeniellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F10EAA55FFA6FF8FA" box="[1218,1436,1836,1861]" italics="true" pageId="12" pageNumber="213">Pseudochlaeniellus</emphasis>
Jeannel, 1949
</taxonomicName>
, eight to sg. 
<taxonomicName id="4C4638C7F3087F6F15E8A53CFD09F8D7" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" box="[448,762,1871,1896]" class="Insecta" family="Carabidae" genus="Chlaeniostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F15E8A53CFD9EF8D7" box="[448,621,1871,1896]" italics="true" pageId="12" pageNumber="213">Chlaeniostenus</emphasis>
Bates, 1892
</taxonomicName>
, four to sg. 
<taxonomicName id="4C4638C7F3087F6F1756A53CFAD9F8D7" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" box="[894,1322,1871,1896]" class="Insecta" family="Carabidae" genus="Amblygenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F1756A53CFBFCF8D7" box="[894,1039,1871,1896]" italics="true" pageId="12" pageNumber="213">Amblygenius</emphasis>
LaFerté-Sénectère, 1851
</taxonomicName>
, six to the nominal genus, two to sg. 
<taxonomicName id="4C4638C7F3087F6F15FAA507FCF3F833" authority="Kolbe, 1889" authorityName="Kolbe" authorityYear="1889" box="[466,768,1908,1933]" class="Insecta" family="Carabidae" genus="Leptorembus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F15FAA507FD90F832" box="[466,611,1908,1933]" italics="true" pageId="12" pageNumber="213">Leptorembus</emphasis>
Kolbe, 1889
</taxonomicName>
, one to sg. 
<taxonomicName id="4C4638C7F3087F6F17A7A507FAF2F833" authority="Kuntzen, 1919" authorityName="Kuntzen" authorityYear="1919" box="[911,1281,1908,1933]" class="Insecta" family="Carabidae" genus="Macrochlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F17A7A507FBB9F832" box="[911,1098,1908,1933]" italics="true" pageId="12" pageNumber="213">Macrochlaenites</emphasis>
Kuntzen, 1919
</taxonomicName>
, seven to sg. 
<taxonomicName id="4C4638C7F3087F6F14BFA5E4FD81F80F" authority="LaFerte-Senectere, 1851" authorityName="LaFerte-Senectere" authorityYear="1851" box="[151,626,1943,1968]" class="Insecta" family="Carabidae" genus="Homalolachnus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F14BFA5E4FEBBF80F" box="[151,328,1943,1968]" italics="true" pageId="12" pageNumber="213">Homalolachnus</emphasis>
LaFerté-Sénectère, 1851
</taxonomicName>
, one to sg. 
<taxonomicName id="4C4638C7F3087F6F1723A5E4FBB9F80F" authority="Peringuey, 1896" authorityName="Peringuey" authorityYear="1896" box="[779,1098,1943,1968]" class="Insecta" family="Carabidae" genus="Pleroticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F1723A5E4FC8EF80F" box="[779,893,1943,1968]" italics="true" pageId="12" pageNumber="213">Pleroticus</emphasis>
<bibRefCitation id="EFD73EB5F3087F6F17A3A5E4FBB9F80F" author="Peringuey" box="[907,1098,1943,1968]" pageId="12" pageNumber="213" refString="Peringuey, L. (1896) A descriptive catalogue of the Coleoptera of South Africa. Part II. Carabidae. Transactions of the South African Philosophical Society, 7, 125 - 623, pls. III - X." type="journal article" year="1896">Péringuey, 1896</bibRefCitation>
</taxonomicName>
, two to sg. 
<taxonomicName id="4C4638C7F3087F6F10CCA5E4FEAFF86A" authority="Basilewsky, 1965" authorityName="Basilewsky" authorityYear="1965" class="Insecta" family="Carabidae" genus="Paracallistoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F10CCA5E4FA68F80F" box="[1252,1435,1943,1968]" italics="true" pageId="12" pageNumber="213">Paracallistoides</emphasis>
Basilewsky, 1965
</taxonomicName>
and five to sg. 
<taxonomicName id="4C4638C7F3087F6F1621A5CFFC9EF86A" authority="Motschulsky, 1865" authorityName="Motschulsky" authorityYear="1865" box="[521,877,1980,2005]" class="Insecta" family="Carabidae" genus="Callistoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B9329F56F3087F6F1621A5CFFD7CF86A" box="[521,655,1980,2005]" italics="true" pageId="12" pageNumber="213">Callistoides</emphasis>
Motschulsky, 1865
</taxonomicName>
.
</paragraph>
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