A revision of the genus Gondwanoscurus Jezek (Diptera: Psychodidae) Author Curler, Gregory R. text Zootaxa 2009 2169 21 34 journal article 10.5281/zenodo.189174 080a1b52-dc1f-4791-b103-4f61f4081b90 1175-5326 189174 Gondwanoscurus praecipuus (Quate) ( Figs. 22 –25) Telmatoscopus praecipuus Quate, 1962 a, 4: 11 (original description); Duckhouse, 1973 , 232 (Oriental catalog). Diagnosis. Adult: Male antennae: flagellomeres 1– 2 each bearing three lateral spines, flagellomere 3 bearing four lateral spines; ascoids each with four digitiform branches, arranged in multiple irregular rows around each node. Male terminalia: gonostyli bifurcate, with medial rami arcuate; parameral sheath constricted medially, expanded apically, not wider than distance between gonocoxal bases; basiphallus laterally compressed basally, bifid apically, with branches of "Y" fused medially, spatulate; distiphallus comprised of paired, sinuous sclerites, articulated with basiphallus basal to "Y"; cercopodia curved approximately 45° at mid-length. Redescription. Adult Male ( Figs. 22 –25): Measurements, (N = 2) head width 0.5 mm (0.49–0.51), head length 0.5 mm (0.49–0.51), wing length 2.78 mm (2.75–2.80), wing width 1.85 mm , palpomere proportion: 1–1.7–1.6–2.3. Head: frontal scar patch subquadrate anteriorly, bilobed posteriorly, without median spur; antennae with scape approximately 2.5 times longer than wide; flagellomere nodes not exserted laterally; flagellomeres 1– 2 each bearing three lateral spines, flagellomere 3 bearing four lateral spines, spines with apices blunt, rounded; ascoids each with four digitiform branches, arranged in multiple irregular rows around each node. Wing: broadly rounded, slightly longer than wide; medial fork basal to radial fork, both arising basal to apex of CuA2; base of M2 incomplete. Terminalia: gonocoxites approximately 1.25 times longer than wide, widely separated at base; gonostyli bifurcate; lateral rami stout, tapered, with acute apices; medial rami of gonostyli arising at about mid-length on medial surface of lateral rami, slender, arcuate, with rounded apices; parameral sheath constricted medially, expanded apically, not wider than distance between gonocoxal bases; basiphallus laterally compressed basally, bifid apically, with branches of "Y" fused medially, spatulate; distiphallus comprised of paired, sinuous sclerites, articulated with basiphallus basal to "Y"; cercopodia curved approximately 45° at mid-length; retinacula spathulate, numbering 7. Adult Female : Unknown. Type material. Holotype [adult male]: NORTH BORNEO [ MALAYSIA , SABAH ]. Tenompok, approximately 50 km E of Jesselton, 18.x.1958 , coll. L. W. Quate. Type deposited at BPBM . Paratype : same locality as holotype , 20.x.1958 [ 1 adult male, partial specimen], coll. L. W. Quate. Paratype deposited at BPBM . Distribution. Known only from the type locality. Remarks. Adult males of G. praecipuus are larger than males of all other described Gondwanoscurus species, and can be readily distinguished by the blunt spines on antennal flagellomeres 1– 3, and the gonostyli with arcuate medial rami. Males of G. praecipuus are most similar to those of G. e x i m i u s in the structure of the antennae, and broadly rounded wings. Two female specimens lacking collection data and labeled only as " praecipuus ?" were examined with the holotype . It is impossible to positively associate these females with the male of G. praecipuus , but the terminalia are similar to females of G. ejundicus , confirming that they are Gondwanoscurus .