Two new species of Clito from South America and a revision of the Clito littera group (Lepidoptera: Hesperiidae: Pyrginae) Author Grishin, Nick V. text Zootaxa 2014 3861 3 231 248 journal article 10.11646/zootaxa.3861.3.2 7beda7dd-c67d-44f3-9e62-cf4500db236c 1175-5326 252285 0F6451EC-F2A1-4EC8-8C86-CB0A4EA73AB9 C. anda Evans 1953 , new status ( Figs. 11–14 , 26 l–o, 27 part, 28 part) C. anda differs most prominently from both C. littera and C. nebulosa in the cucullus being strongly bulged distad at the bend ( Figs. 26 l–o), while the ampulla is similar to C. littera . Wing patterns of these three species also amply differ, as stated in the key below and indicated on Fig. 27 . Due to these differences in genitalia and facies, it seems more reasonable to treat these taxa as distinct biological species rather than subspecies. In researching this question, a difficulty was encountered. Both valvae of C. anda [holo] type , glued to the carton card by Evans, have been damaged and the cuculli broken off. Apparently, the damage happened after the sketches had been made from them by Evans. These sketches: one pinned in the BMNH collection, the other one published in the book ( Evans 1953 )—while being similar to each other, differ in slight details ( Figs. 26 n–o). Both show very prominent distal expansion of the cucullus (bulge). On the sketch from the collection, this bulge is less expressed, more consistent with a more accurate drawing by Austin (2000) of C. anda specimen from Brazil : Rondônia ( Fig. 26 l). This collection sketch shows a more prominent rise in the ampulla, also more similar to the Austin drawing, and, more importantly, more similar to the holotype valva ( Fig. 26 m). It appears that the collection sketch is more accurate. Importantly, since the cuculli were broken off the holotype genitalia mount, both of these sketches made from the holotype (TL: French Guiana , the sole [para] type was not dissected, see Figs. 13–14 ) and Austin drawing from a Rondônia specimen show the distal bulge of the cucullus in C. anda , thus setting this species apart from both C. littera and C. nebulosa , the two species without this distal expansion and the cucullus narrowing gradually towards the dorsally pointed tip. After Evan's work, two more Clito species with apparent phylogenetic affinities to C. littera have been discovered.