Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr Author Spiegel, Didier Van Den Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be text Zoosystema 2007 29 3 417 456 journal article 10.5281/zenodo.5393630 1638-9387 5393630 Glyphiulus paracostulifer n. sp. ( Figs 22-24 ) TYPE MATERIAL . — China . Guizhou Prov. , Qianxy , Hong Lin Town , Lao Hu Dong Cave , 21.XI.2003 , leg. L. Latella & S. Meggiorini , holotype ♂ ( MNHN GA 052 ) ; paratypes 1 ♀ ( MNHN GA 052 ) ; 1 ♀ ( ZMUM ) ; 1 ♀ ( MCSNV / SEM ) . FIG. 20. — Glyphiulus percostulifer n. sp. , paratype ♀: A , anterior part of body, lateral view; B , midbody segment section; C , midbody segments, lateral view; D , same, dorsal view; E , caudal body end, lateral view; F , same, ventral view. Scale bars: A-E, 0.2 mm; F, 0.1 mm. ETYMOLOGY. — To emphasize the close resemblance to G. costulifer n. sp. DIAGNOSIS. — Differs from congeners in the relatively large size, the completely unpigmented tegument and ocelli, the undivided tergal crests, coupled with certain details of leg and gonopod structure (see also key below). FIG. 21. — Glyphiulus percostulifer n. sp. , paratype ♂: A , antenna; B , gnathochilarium, ventral view; C , leg 1, caudal view; D , leg 2, caudal view; E , leg 3; F , leg 13; G , anterior gonopods, caudal view; H , right posterior gonopod, frontal view. Scale bar: 0.2 mm. DESCRIPTION Holotype with 65p+1a+T, about 43 mm long and 2.0 mm wide; adult paratypes with 52p+2a+T, 55p+3a+T or 57p+1a+T, all about 36 mm long and 2.0 mm wide. Coloration rather uniformly pale yellowish to brownish, only some ozoporiferous cones a little darker; ocelli unpigmented, a few barely discernible per eye patch. Gnathochilarium (mentum divided, n = 1), postcollar constriction ( Fig. 22B ), segment’s texture and cross-section ( Fig. 22 A-E), most of telson ( Fig. 22E, F ), ventral flaps of male segment 7, male legs 2 and 3 ( Fig. 24C, D ), posterior gonopods ( Fig. 24G ) generally as in G. subcostulifer n. sp. , but antennae slightly more elongate and less rich in sensilla ( Figs 22A ; 23D ; 24A ); collum ( Fig. 22A, B ) with a nearly typical carinotaxy formula (I-III+4c+5a+pc+ma+pc+5a+4c+III- I), crests evident; all tergal crests ( Figs 22 A-E; 23A) on subsequent body segments higher, especially so until about segment 9 or 10, virtually not divided (2+I/i+3+I/i+2), ozoporiferous cones only barely lower than wide; epiproct with an evident dorsal rib ( Fig. 22E ); legs about as long as body diameter ( Fig. 23A, B ), at most with only a vestigial spine at claw base ( Figs 23C ; 24E ); male leg 1 with reduced 5-segmented telopodites and large paramedian coxal processes ( Fig. 24B ); each anterior gonopod plate with a high distomedial process ( Fig. 24F ). FIG. 22.— Glyphiulus paracostulifer n.sp. , paratype ♀: A , anterior part of body,lateral view; B , same,dorsal view; C , midbody segments, lateral view; D , same, dorsal view; E , caudal body end, lateral view; F , same, ventral view. Scale bars: 0.5 mm. FIG. 23. — Glyphiulus paracostulifer n. sp. , paratype ♀: A , midbody segment section; B , midbody leg; C , claw; D , antennal sensilla. Scale bars: A, 0.5 mm; B, 0.2 mm; C, 0.1 mm; D, 0.05 mm. REMARKS Based on several troglomorphic traits, such as the unpigmented body and ocelli, as well as the somewhat elongated antennae and legs, this species seems to be troglobitic.