Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr Author Spiegel, Didier Van Den Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be text Zoosystema 2007 29 3 417 456 journal article 10.5281/zenodo.5393630 1638-9387 5393630 Glyphiulus costulifer n. sp. ( Figs 14-16 ) TYPE MATERIAL . — Laos . Luang Prabang Prov. , Nong Kiaw ( Muang Ngoy ), Tham Pha Kouang Cave , 102.68°E , 20.72°N , 29.XII.2000 , leg. A. Bedos & L. Deharveng (LAO-099), holotype ♂ ( MNHN GA 049 ) ; paratypes 3 ♂♂ , 2 ♀♀ ( MNHN GA 049 ) ; 1 ♂ ( SEM ) ; 1 ♂ , 1 ♀ ( ZMUM ) ; 1 ♂ , 1 ♀ ( ZMUC ) ; 1 ♀ ( NMNHS ) . ETYMOLOGY. — To emphasize the normally carinate collum. DIAGNOSIS. — Differs from congeners by the nearly typical carinotaxy pattern of the collum, coupled with highly unusual traces of both ventrolateral crests, the very little modified male coxae 1, the long but bare flagellum of the posterior gonopods, etc. (see also key below). DESCRIPTION Holotype with 56p+1a+ T , about 28 mm long and 1.8 mm wide; paratypes ( ♂ , ♀ ) with 48-62p+3-1a+ T , length 23-35 mm , width 1.5-1.9 mm . Coloration light yellow-brown to red-brown, tergal crests often dark castaneous brown. Greyish to red-brown, subtriangular eye patches with 7-9 ocelli each. Antennae usual ( Figs 14A ; 16A ), gnathochilarium (mentum divided, n = 3) oligotrichous ( Fig. 16B ). Head width = segment 2 = 6 <collum = 7(8)> segment 3 = 5 <7 <8 <9 <10 <11 = midbody segments; body gradually tapering toward telson. Postcollar constriction evident ( Fig. 14B ). Collum almost typically crested, only median crest (ma) unusually abbreviated; carinotaxy pattern typical of the granulatus -group (I-VI+7a+pc+ma+pc+7a+VI- I) ( Fig. 14A, B ). Subsequent metaterga evidently crested as well ( Fig. 14 A-E).Carinotaxy formula 2+I/ i+3/3+I/i+2, i.e. all dorsal crests subdivided ( Fig. 14C, D ), while both ventrolateral crests undivided, barely traceable as ribbons of micropunctate/shagreened texture ( Fig. 15B ). Ozoporiferous tubercles round, only slightly wider than high ( Fig. 15A, B ). Tegument delicately alveolate-areolate, dull throughout. Fine longitudinal striations in front of stricture between pro- and metazona ventrolaterally, laterally and dorsally, remaining surface of prozona very delicately shagreened. Segments round in crosssection ( Fig. 15A ). Epiproct with a distinct, humplike ridge dorsally in distal 3/4 extent ( Fig. 14E ), in ventral view rounded and subtriangular in caudal part ( Fig. 14F ). Paraprocts flattened ventrally, with only a shallow but evident submarginal depression beside caudal edge, each with a row of about a dozen setae at medial margin ( Fig. 14F ). Hypoproct sublanceolate, very delicately and broadly emarginate caudomedially ( Fig. 14F ). Ventral flaps behind gonopod opening on male segment 7 distinguishable as a low but clear transverse ridge ( Fig. 15C ). Legs long, those on midbody segments about as long as segment diameter ( Fig. 14C ). Claw with a short but evident accessory spine at base ( Fig. 16 C-F). Male legs 1 highly characteristic ( Fig. 16C ) in showing only slightly modified coxae with two low, paramedian, contiguous cones; telopodites 5-segmented, normal. Male legs 2 and 3 modified as usual ( Fig. 16D, E ). Anterior gonopods ( Figs 15D ; 16G ) not notched, paramedian processes (d) high, telopodites evidently curved. Posterior gonopods ( Figs 15E, F ; 16H ) highly compact, typical of the group, flagella (f) high and bare. REMARKS Due to the nearly unmodified male legs 1, this species is probably among the basalmost in the genus. In the absence of any troglomorphic traits, this species can only be considered troglophilic at most. This cave is known to support another congener, G. bedosae Golovatch, Geoffroy, Mauriès & Van den Spiegel, 2007 (see Golovatch et al. 2007 ). Interestingly, both species share the same, rather conspicuous structure of the telson.