The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist Author Freese, Edwin L. 33493 “ S ” Avenue Adel, Iowa 50003 Author Veal, Doug A. 168 Thompson Drive SE Cedar Rapids, Iowa 52403 Author Lago, Paul K. Biology Department, University of Mississippi University, Mississippi 38677 text Insecta Mundi 2020 2020-08-28 2020 787 1 83 journal article 10.5281/zenodo.5353792 1942-1354 5353792 88D45C5E-0E49-4F02-87A4-C4FA55A83B2F Phyllophaga vehemens ( Horn, 1887 ) Travis 1934 ; ISIC; IWUC; UMRC; ELFC; DAVC; PKLC. Collection dates: April–July. Collected at light. Travis (1934) reported 434 specimens collected from Iowa indicating this species was infrequent. County records ( Figure 150 ): Appanoose, Clayton, Dallas, Fremont, Harrison, Iowa , Keokuk, Lee, Louisa, Mahaska, Mills, Monona, Ringgold, Story, Wapello.