Revision of the Empis subgenus Enoplempis Bigot, east of the Rocky Mountains (Diptera: Empididae) Author Sinclair, Bradley J. Author Brooks, Scott E. Author Cumming, Jeffrey M. text Zootaxa 2013 3736 5 401 456 journal article 10.11646/zootaxa.3736.5.1 495ee224-dd4f-4c96-aa06-094b23e135a1 1175-5326 248431 2A05A2A2-CF49-4585-A75D-7086B9DDD7B3 Key to males of E . ( Enoplempis ) east of the Rocky Mountains 1 Hind femur and tibia without processes, setal cones or distinct combs of setae ( Figs 3 A, E, 6E, 12E, 19A, D, E); hind tro- chanter with ( Figs 3 C, E, 9D, 12D, 16B, 19C) or without modified setae or processes................................ 2 - Hind femur and/or tibia with subapical/subbasal processes, setal cones or distinct combs of setae ( Figs 6 A, D, 9A, E, 12A, C, 16A, C, 23A); hind trochanter lacking modified setae........................................................ 11 2 Hind trochanter bearing modified setae or processes ( Figs 3 C, 9D, 12D, 16B, 19C)................................. 3 - Hind trochanter unmodified, setae unmodified............................................................. 8 3 Thoracic pleura yellow, in contrast to dark, greyish scutum. Hind trochanter with 5–8 stout setae widespread, not confined to process or arising from single spot ( Fig. 3 C)............................................... E. ( En. ) amytis Walker - Thoracic pleura greyish, concolorous with scutum ( Fig. 3 D). Hind trochanter with stout setae confined to process or arising from single spot; if setae more widespread, then greater than 8 present ( Figs 9 D, 12D, 16B, 19C)..................... 4 4 Hind tibia with numerous rows of long slender dorsal setae, longer than width of tibia ( Figs 3 E, 12E).................. 5 - Hind tibia with only widely spaced setae, at most only slightly longer than width of tibia ( Figs 9 C, 16B, 19B)........... 6 5 Setae of hind trochanter confined to distinct spot or short tubular process ( Fig. 12 D). Apex of phallus expanded, funnel-shaped ( Fig. 15 A)............................................................................ E. ( En. ) nuda Loew - Setae of hind trochanter widespread, not confined to distinct spot or short process ( Fig. 3 E). Apex of phallus slender, tapered to narrow apex ( Fig. 4 B)................................................... E. ( En. ) appalachicola Sinclair sp . nov . 6 Setae of hind trochanter widespread, not confined to distinct spot or process ( Fig. 19 C). Wing length less than 6 mm ....................................................................................... E. ( En. ) stenoptera Loew - Setae of hind trochanter confined to distinct spot or process ( Figs 9 C, 16B). Wing length greater than 6 mm ............. 7 7 Setae of hind trochanter confined to distinct tubular process ( Fig. 9 D). Hypandrium 0.5× length of epandrium ( Fig. 10 A), apex tapered and rounded; setae present on membrane anterior to base of hypandrium.............. E. ( En. ) gulosa Coquillett - Setae of hind trochanter confined to distinct spot, not raised on tubercle ( Fig. 16 B). Hypandrium short, about one-third length of epandrium ( Fig. 17 A), apex notched; setae absent from membrane anterior to hypandrium................................................................................................... E. ( En. ) penicillata Brooks sp . nov . 8 Scutum with three distinct narrow vittae. Wing with clouding at base of R4 and apex of discal cell (Fig. 2B). Hind femur robust with anterior row of ventral spine-like setae ( Fig. 19 D)................................. E. ( En. ) tridentata Coquillett - Scutum with 2 or 4 indistinct broad vittae. Wing lacking clouding. Hind femur more slender, lacking ventral row of stout setae ( Figs 6 E, 19A, E)..................................................................................... 9 9 Hind femur and tibia stout, with some spine-like setae; hind femur lacking rows of very slender, pale ventral setae ( Fig. 6 E). Epandrium quadrate, not tapered posteriorly; hypandrium prolonged posteriorly, 0.5× length of epandrium ( Fig. 8 A)........................................................................................ E. ( En. ) enodis Melander - Hind femur and tibia long and slender, lacking spine-like setae; hind femur with rows of very slender, pale ventral setae ( Figs 19 A, E). Epandrium tapered posteriorly; hypandrium short, less than half length of epandrium ( Figs 20 A, 21B)......... 10 10 Apex of epandrium greatly narrowed, less than half width of base of epandrium ( Fig. 20 A). Posterior margin of hypandrium truncate, not produced posteriorly; bearing long setae greater than length of hypandrium. Cercus broad apically with numerous sinuous setae..................................................................... E. ( En. ) snoddyi Steyskal - Apex of epandrium not greatly narrowed, more than half width of base of epandrium ( Fig. 21 B). Posterior margin of hypandrium produced, rounded; bearing short setae less than length of hypandrium. Cercus greatly narrowed on posterior third, bearing numerous straight, stout setae.......................................... E. ( En. ) vockerothi Cumming sp . nov . 11 Hind femur and tibia with dense rows of preapical setae, lacking highly modified processes ( Figs 6 D, 12C, 16A)....... 12 - Hind femur and tibia with highly modified processes ( Figs 6 A, 9A, E, 12A, 16C, 23A)............................ 14 12 Hind femur lacking preapical row of dense setae, only hind tibia with dense setae ( Fig. 6 D). Epandrium quadrate with angular margins and fused anterodorsally to cercus ( Fig. 7 B).................................. E. ( En. ) ctenocnema Melander - Hind femur with preapical posteroventral and/or anteroventral row of dense setae, similar to setae of tibia ( Figs 12 C, 16A). Epandrium more rounded apically, not fused to cercus ( Figs 13 B, 15B).......................................... 13 13 Hind femur with anteroventral preapical row of stout setae ( Fig. 12 C). Base of hind tibia with pair of antero- and posteroventral rows separated by clear gap ( Fig. 12 C). Hypandrium with setae longer than length of sclerite ( Fig. 13 B)........................................................................................... E. ( En. ) nodipoplitea Steyskal - Hind femur with antero- and posteroventral preapical rows of stout setae ( Fig. 16 A). Base of hind tibia with rows of setae dense, not separated by clear gap ( Fig. 16 A). Hypandrium with setae shorter than length of sclerite ( Fig. 15 B)...................................................................................... E. ( En. ) pectinata Sinclair sp . nov . 14 Thoracic pleura primarily yellow....................................................................... 15 - Thoracic pleura primarily greyish, not predominantly yellow................................................. 16 15 Base of hind tibia with pair of slender ventral processes with narrow brush of black apical setae ( Figs 12 A, B). Hypandrium clothed in long, dense setae ( Fig. 13 A). Phallus without lateral projections, only slightly tapered towards apex ( Fig. 13 A).......................................................................... E . ( En. ) montywoodi Brooks sp . nov . - Base of hind tibia with pair of blunt ventral processes, posteroventral process distinctly tufted, anteroventral process anvilshaped ( Figs 16 C, D, E). Hypandrium with setae shorter, confined to base ( Fig. 17 B). Phallus flanked by stout lateral pointed projections, medial portion denticulate and slender apically ( Fig. 17 B)............. E . ( En. ) prodigiosa Cumming sp . nov . 16 Hind tibia with setae subequal to or shorter than width of segment ( Figs 6 A, 23A). Hind femur with tufted preapical anteroventral process ( Figs 6 B, 23B)............................................................................ 17 - Hind tibia with setae longer than width of segment ( Fig. 9 E). Hind femur lacking tufted preapical anteroventral process ( Figs 9 A, B, F, G)........................................................................................ 18 17 Base of hind tibia geniculate and highly excavated, fitting posteroventral processes of femur ( Figs 6 B, C); apical half of phallus straight, directed dorsally ( Fig. 7 A)............................................... E. ( En. ) arthritica Melander - Base of hind tibia not geniculate, with pair of tufted processes ( Figs 23 B, C); apical half of phallus strongly arched, directed anterodorsally ( Fig. 24 )....................................................... E. ( En. ) volsella Sinclair sp . nov . 18 Base of hind tibia with thumb-like process clothed with short, fine setae ( Fig. 9 A). Hind femur with hooked, posteroventral preapical process longer than width of femur ( Figs 9 A, B)............................... E. ( En. ) gladiator Melander - Base of hind tibia with bare, elongate, pointed triangular process ( Figs 9 F, G). Hind femur with triangular, posteroventral preapical process shorter than width of femur ( Figs 9 F, G)............................... E. ( En. ) loripedis Coquillett