Revision of New World species of Roederiodes Coquillett (Diptera: Empididae: Clinocerinae) Author Sinclair, Bradley J. Canadian Food Inspection Agency, K. W. Neatby Bldg., C. E. F., 960 Carling Ave., Ottawa, ON, Canada K 1 A 0 C 6 & Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Bldg., C. E. F., 960 Carling Ave., Ottawa, ON, Canada K 1 A 0 C 6 text Zootaxa 2023 2023-06-12 5301 3 336 364 http://dx.doi.org/10.11646/zootaxa.5301.3.2 journal article 10.11646/zootaxa.5301.3.2 1175-5326 8030504 9451FCC7-EA59-4ACE-B45F-8B012B19D8AA Roederiodes browni sp. nov. ( Fig. 16 , 31 ) urn:lsid:zoobank.org:act: 31B6085A-038E-4C59-BD84-3F1925456469 Type material. HOLOTYPE ♁, labelled: “ COSTA RICA : 1500m / Monteverde , Finca / Canada [ 10°18′58″N 84°48′28″W ], 5.VI.1988 / BV Brown, native for.”; “ CNC DIPTERA / #103484”; “ Barcode of Life / DNA voucher specimen/ Smple CNC DIPTERA 103484 / BOLD Proc. ID: CNCDD185-11 [green label]”; “ HOLOTYPE / Roederiodes / browni Sinclair [red label]” (barcoded, CNC ). Recognition. This species is similar in colour to the other Central American species and can be distinguished by the quadrate clasping cercus, not longer than height of epandrium, slightly projecting anteriorly and posterior margin narrowly expanded. Description. Wing length 2.3 mm . Male. Face and gena without bluish pubescence. Ocellar setae divergent, slightly shorter than pprn; vertical seta as long as ocellar seta. Antenna with short stylus, slightly longer than postpedicel; postpedicel ovate, shortly prolonged apically. Length of proboscis slightly shorter than head height. Pleura pale brown without bluish pruinescence; prescutellar depression without pruinescence, postalar ridge paler than scutum; several acr, very short, anterior to second pair of dc; 5 dc, long, prescutellar dc longest; 1 pprn; 1 presut spal; 2 npl; 0 psut spal; 1 pal; 2 sctl, longer than prescutellar dc and numerous setulae on disc and margin, outer marginal setulae one-half length of apical seta; laterotergite with several pale setae; antepronotum with several setulae; lower proepisternum with pale setae. Wing infuscate; M 1 and M 2 separated by short crossvein, shorter than crossvein r-m. Legs brown, with coxae pale brown, concolourous with pleura; coxae without bluish pruinescence. Hind tibia with weak ventral setae. Abdominal tergites brown, darker than pleura, without bluish pruinescence; bearing short pale setae. Terminalia ( Fig. 16 ): Hypandrium slightly longer than epandrium, apex rounded; without setae. Phallus straight; membranous distiphallus with pair of elongate lateral extensions, bearing short, apical sclerotized plates. Cercal plate with 2 long setae; clasping cercus quadrate, not longer than height of epandrium; anterior margin slightly prolonged; posterior margin narrowly expanded; stout, thickened setae along inner dorsal margin; longer, finer setae on inner subdorsal and anterior margins. Female. Unknown. Etymology. This species is named in honour of Brian V. Brown, curator at the Natural History Museum of Los Angeles County, who collected the holotype . Distribution. This new species is known only from the type locality at Estación Biológica Monteverde, in Costa Rica ( Fig. 31 ). Remarks. This is one of three species collected from a small creek (Máqùina) below the research building at Estación Biológica Monteverde. This is a private biological research station with ca. 120 hectares of secondary and primary cloud forest located on the Pacific slope of the Tilarian Mountain Range. Each species appears to be collected at different periods of the year. This species was collected in June (start of wet season), whereas R. woodi sp. nov. was collected in February and March (dry season) and R. costaricensis sp. nov. in August (wet season). Future collections throughout the year are required to investigate further the apparent temporal separation of these species. The barcode obtained from the holotype was highly fragmented with many gaps, so fresh collections are required to confirm the barcode identification of this species.