| Characters | States | |
|---|---|---|
| 1. | DS (dorsal scutum), main outline in dorsal view |
(0) Beta, abdominal part size comparable to carapace, steadily growing wider posteriorly, with broad convex posterior margin (typical of
|
| 2. | Posterior margin of DS, shape | (0) convex; (1) straight; (2) concave. |
| 3. | Carapace outline, sexual dimorphism | (0) same size in both sexes; (1) greatly enlarged in males, groove projecting backwards, squeezing scutal area I. |
| 4. | Ocularium, structure | (0) high, coarsely granular, slanted forward; (1) low, may even have a median depression. |
| 5. | Placement of the eyes on two separate eyeballs (absence of ocularium) | (0) eyes placed on a common ocularium; (1) 2 separate eyes. |
| 6. | Frontal hump of carapace | (0) absent; (1) present. |
| 7. | Scutal area I, presence of median longitudinal groove | (0) absent, area I entire; (1) present, area I divided into left and right halves; (2) secondary effacement; (3) Goniosoma-like (area II invading area I). |
| 8. | Scutal area III, paired armature | (0) unarmed; (1) with paramedian pair of tubercles or spines; (2) high spines, partially fused. |
| 9. | Median spiniform apophysis of free tergite III, shape | (0) absent; (1) stout; (2) small. |
| 10. | Median spiniform apophysis of free tergite II, presence | (0) absent; (1) present |
| 11. | Furrow between scutal areas III and IV, degree of effacement | (0) furrow complete; (1) furrow effaced in the middle; (2) furrow entirely effaced. |
| 12. | Furrow between areas III and IV, shape | (0) straight; (1) clearly recurved (pointing forwards); (2) gently procurved (pointing backwards); (3) barely distinct, sinuous (formed by two, joined, Cshaped furrows) clearly marked, sinuous; (4) entirely effaced. |
| 13. |
Lateral margins of dorsal scutum, armature (character inactive, see
|
(0) finely granular or without special granulation arranged in a row; (1) with longitudinal row of light-colored, large tubercles, coalescent and flattened; (2) with marginal row of well-spaced, light-colored tubercles. |
| 14. | Stigmatic area, posterior border | (0) straight or only very slightly concave; (1) deeply concave. |
| 15. | Basichelicerite, structure | (0) short, unarmed; (1) pre-bulla very long, powerfully armed; (2) bulla long, robust, with marginal armature; (3) bulla of male attenuate; (4) long and very thick. |
| 16. | Posterior margin of cheliceral bulla, ornamentation | (0) smooth; (1) with several teeth. |
| 17. | Cheliceral hand of male, sexual dimorphism | (0) present, cheliceral hand balloon-like, swollen; (1) absent, cheliceral hand similar in both sexes; (2) intermediate, chelicera of male a little larger. |
| 18. | Cheliceral fingers, elaborate dentition, presence | (0) absent, teeth reduced and subequal; (1) present, teeth conform to varied categories, fingers often twisted and bent. |
| 19. | Pedipalpal trochanter, modification for locking | (0) without modifications; (1) with ventro-basal apophysis interlocking against cavity in coxa I. |
| 20. | Pedipalpal femur, shape | (0) cylindrical; (1) compressed, slightly concave mesally, with ventral row of numerous small spines coalesced into a flap; (2) compressed, with ventral row of independent small spines; (3) slightly compressed, accommodated wrapping chelicerae. |
| 21. | Pedipalpal femur, disto-mesal setiferous tubercle | (0) absent; (1) present. |
| Characters | States | |
|---|---|---|
| 22. | Pedipalpal femur, ventral row of spines | (0) absent; (1) numerous short teeth; (2) few stout spines; (3) one basal larger and a few smaller in the middle. |
| 23. | Pedipalpal femur, ventro-basal cluster of setiferous tubercles | (0) only one, clearly ventral, outside the 2 rows of ventro-ectal and ventro-mesal setiferous tubercles; (1) a cluster of 2 divergent setiferous tubercles; (2) none. |
| 24. | Pedipalpal tibia, shape | (0) semi-cylindrical, without flap; (1) strongly depressed and concave, all setae fused forming a marginal flap. |
| 25. | Pedipalpal tibia, ventro-ectal setiferous tubercles, size relationship | (0) subequal; (1) penultimate or last much larger. |
| 26. | Pedipalpal tarsus, shape | (0) large, subrectangular, with normal claw; (1) small, subtriangular with claw much reduced. |
| 27. | Patella and tibia of pedipalp, granulation of dorsal surface | (0) finely granular; (1) with coarse tuberculation. |
| 28. | Pedipalp, general size compared to body | (0) as long as scutum; (1) extremely elongate and thin; (2) robust, clearly larger than scutum; (3) stunted, shorter than scutum and thin; (4) not comparable, cosmetiform. |
| 29. | Tarsus I, basal articles fused forming a spindle | (0) absent; (1) present. |
| 30. | Relative thickness of legs I–IV | (0) all subequal; (1) III–IV clearly stronger than I–II; (2) legs getting steadily stronger from I to IV; (3) only leg IV much stouter than the others in the males. |
| 31. | Coxa IV, dorso-apical apophysis only in male | (0) absent; (1) present. |
| 32. | Coxa IV of male, armature | 0 weakly developed, armed with small spine or unarmed; (1) very robust, hornlike apophysis. |
| 33. | Coxa IV of male, position relative to dorsal scutum in dorsal view | (0) concealed under DS; (1) clearly surpassing DS. |
| 34. | Trochanter IV of male, retro-distal region, armature | (0) unarmed; (1) stout recurved hook; (2) short, spiniform apophysis; (3) stout, procurved hook. |
| 35. | Femur IV of male, structure | (0) elongate, thin, cylindrical, substraight; (1) short, slightly sinuous, thickened; (2) moderately long, only slightly incrassate sinuous. |
| 36. | Femur IV, sexual dimorphism | (0) absent; (1) clearly marked. |
| 37. | Tibia IV, sexual dimorphism | (0) absent; (1) clearly marked |
| 38. | Distitarsus of leg I, segmentation | (0) with 2 articles; (1) with 3 articles. |
| 39. | Tarsal claw, legs 3–4, structure | (0) median prong, with lateral secondary prongs; (1) paired claws. |
| 40. | Tarsal process | (0) absent; (1) present, similar to claws; (2) present, very thick. |
| 41. | Truncus penis, presence of musculature | (0) present, one muscle along the shaft; (1) absent, gland moved by hydraulic pressure. |
| 42. | Pars distalis of truncus penis, structure | (0) Distal region of truncus bears MS, but it is not clear-cut defined, being continuous with pars basalis; (1) A Ventral plate (VP) is present in the form of a lamina parva, separated from the base by a waist; (2) completely defined as a flattened subrectangular VP, clearly separated from truncus; (3) Truncus uniform, topped by a tongue-shaped setigerous plate, homology with macrosetae of other Laniatores is uncertain; (4) VP well-defined, immensely developed, complex-shaped, separated from truncus by a neck. |
| 43. | Staff-like break on distal portion of truncus, presence | (0) absent; (1) present. |
| 44. | Division of distal setigerous region of truncus | (0) complex system of plates detached from truncus; (1) gradual narrowing of apex without distinction of a podium or a ventral plate; (2) lamina parva wellmarked but extending dorsally, podium absent; (3) distal truncus well-cut as a podium, and VP well-marked. |
| Characters | States | |
|---|---|---|
| 45. | Ventral plate, position in relation to truncus | (0) in the same plan; (1) bent upon ventral plate |
| 46. | Ventral plate length | (0) long, at least twice as long as truncus width; (1) short, as long as truncus width. |
| 47. | Ventral plate, insertion in truncus apex | (0) not sunken into truncus; (1) sunken into truncus. |
| 48. | Lateral borders of pars distalis | (0) without keeling; (1) distorted, thinner, with setae dislodged from lateral to dorso-lateral insertion; (2) flange; (3) flange present but rudimentary only; (4) not comparable because flange region is completely overwhelmed by immense MS C. |
| 49. | Metasarcine spiny sacs, presence | (0) absent; (1) present as a pair of haematodochae latero-subdistally on truncus penis |
| 50. | Ventral plate, overlapping with distal truncus | (0) nor overlapping; (1) truncus reaching about half the length of ventral plate; (2) truncus overlapping very little, only at base |
| 51. | Ampycus-like VP, oval, with deep cleft on distal border, presence | (0) absent; (1) present. |
| 52. | Glans penis dorsal process, presence | (0) absent; (1) present, thumb-like; (2) laminar as a keel. |
| 53. | Glans penis ventral process, presence | (0) absent; (1) gonyleptid process, formed by stem plus flabellum; (2) serrate funnel, arising straight from mid-stylus. |
| 54. | Glans, shape | (0) embodied in the capsula externa; (1) as a haematodocha, relatively free in the apical part of truncus; (2) mounted atop the follis. |
| 55. | Glans complex, size and position in relation to ventral plate | (0) stylus short, atop a short glans; (1) stylus long, in situ surpassing VP because situated on a long podium + glans; (2) stylus very long, atop a short glans, in situ surpassing ventral plate; (3) stylus elongate, upon a long column, surpassing ventral plate; (4) stylus very short, atop a triangular glans. |
| 56. | Sac glans, structure | (0) non-columnar; (1) columnar. |
| 57. | Basal glans sac, structure | (0) formed by amorphous folds; (1) formed by superimposed rings |
| 58. | Stylus, distal portion, shape | (0) cylindrical, or otherwise non-compressed; (1) somewhat compressed, with small serrate crest and ill-defined carena; (2) clearly compressed, with welldeveloped ventral serrate crest and well-marked carena. |
| 59. | Stylus, structure |
(0) cylindrical, with terminal opening; (1) strongly flattened and expanded as a serrate keel, with opening as a slit displaced to dorsal; (2)
|
| 60. | Stylar cap, presence | (0) absent; (1) present. |
| 61. | Ventral plate or ventral surface of penis, microsetae cover | (0) glabrous, microsetae absent; (1) covered by one or two fields or isolated clumps of microsetae of varied shapes. |
| 62. | Median field of scale-bristles, shape | (0) absent; (1) complete; (2) divided in 2 “lungs”; (3) extremely reduced to 2 small flecks. |
| 63. | Median field of scale-bristles, structure | (0) formed by T1; (1) formed by T2 or T2/T3; (2) formed only by T3; (3) formed by T5. |
| 64. |
Lateral fields of scale-bristles (always formed by
|
(0) absent; (1) restricted to corners of ventral plate; (2) prominent, occupying most of ventral plate; (3) two wide stripes almost touching each other in the center; (4) restricted to the area of macrosetae group C. |
| 65. | Macroseta A, position | (0) inserted on lateral border of ventral plate; (1) encircling ventral plate as a girdle. |
| Characters | States | |
|---|---|---|
| 66. | Macroseta A-B, size and arrangement | (0) all fairly elongate arranged as a girdle, B more ventral; (1) lacking entirely; (2) A1–A2 forming slanted row, with B extremely reduced; (3) all relatively sturdy, forming an arch, B a little weaker, always positioned latero-ventralmost in the arch; (4) all a little reduced in size and number, B proximal to A, lateral or ventro-lateral-two A and 1 B forming a triangle, with A lateral in a longitudinal row and B slightly shifted to latero ventral; (5) both A and B inserted at extreme distal end, all robust and long. B lateral to ventro-lateral; (6) Macroseta A forming an evenly spaced row ith C on laterals of ventral plate. |
| 67. | Macroseta A, position | (0) more proximal, with gap between A and C; (1) more distal, adjacent to C. |
| 68. | Macroseta B, presence | (0) absent; (1) present. |
| 69. | Macrosetae C, basic placement | (0) composing with A an outer girdle to glans, outside D; (1) not as a girdle, neither aligned with A, but still dorso-lateral on ventral plate; (2) as a longitudinal lateral row distal on ventral plate. |
| 70. | Macrosetae C, shape |
(0) short, sturdy, at least partly lanceolate; (1) robust, with point lanceolate; (2) extremely long and slender; (3) very long and flattened; (4) robust, “buffalo- horn” (
|
| 71. | Macrosetae D | (0) 1 pair, inserted fully dorsal, at the midway sides of the glans; (1) 1 pair, inserted much more distally on dorsal surface of VP, shifted more to lateral; (2) 1 or 2 pairs, well developed, shifted to distal, but still clearly dorsal; (3) 2 pairs, very small, dorso-basal near the glans; (4) 2 pairs, forming a mid-dorsal comb, located basal to glans. |
| 72. | Macrosetae D, insertion | (0) 1 to 3 pairs fully dorsal, surrounding glans, 1 pair at least as base of glans; (1) only one pair, migrated to latero-dorsal among A-C; (2) one pair dorso- lateral at base of glans; (3) 1 or 2 pairs, fully dorsal, much distal; (4) 1 pair lateral between A and C. |
| 73. | Macrosetae E, position | (0) 2 pairs, entirely on the ventral surface of VP, forming a square; (1) 2 to 4 pairs, strongly reduced, inserted on the latero-distal flange of ventral plate. |
| 74. | Macrosetae E, size | (0) large, size comparable to A, B or C; (1) minute, size much smaller than A, B or C. |
| 75. | Macrosetae, pattern | (0) Pattern A–E not recognizable. Not organized in 5 groups and distributed around the VP surface. Homology obscure; (1) Pattern A–E clearly recognizable within the A, B, C, D, E chaetotaxy. |
| 76. | Tarsal claws of legs III–IV, structure | (0) triaenonychium; (1) double claw. |
| 77. | Tarsal aggregate pores | (0) absent; (1) present. |
| 78. | Row of rounded and large tubercles on lateral margin | (0) lacking; (1) present, coalescent and flattened; (2) present, rounded and well- spaced. |
| 79. | White tubercles on ocularium | (0) lacking; (1) present. |
| 80. | White tubercles on area I | (0) lacking; (1) present. |
| 81. | White tubercles on area II | (0) lacking; (1) present. |
| 82. | White tubercles on lateral margin | (0) lacking; (1) all tubercles; (2) only the largest pair. |
| 83. | Enlarged tubercles or elevation almost at end of lateral margin | (0) lacking; (1) enlarged tubercles; (2) wide elevation. |
| 84. | Free tergite II with one pair of tubercles longer than tergite length | (0) lacking; (1) present. |
| 85. | Free tergite III with one pair of tubercles longer than tergite length | (0) lacking; (1) present. |
| 86. | Lateral margins of areas with white patches | (0) lacking; (1) present. |
| 87. | White patches on posterior region of prosoma | (0) lacking; (1) present. |