Resolving the synonymy and polyphyly of the ‘ Drosophila bakoue species complex’ (Diptera: Drosophilidae: ‘ D. montium species group’) with descriptions of two new species from Madagascar Author Yassin, Amir Author Suwalski, Arnaud Author Raveloson Ravaomanarivo, Lala H. text European Journal of Taxonomy 2019 2019-06-18 532 1 26 journal article 26436 10.5852/ejt.2019.532 fbea3203-a10b-4e4f-b5f3-3d1387bc84c3 3251569 1301752E-3FD5-4F3E-A4F3-6766D18C709A ‘ D. seguyi species complex’ Lachaise, 1971 Figs 1–2 , 3A , 4 , 5 E–F, 6 ‘ D. seguyi species complex’ Lachaise, 1971: 1623 . ‘ D. bakoue species complex’ Rafael, 1984: 179 , syn. nov. Diagnosis Male abdominal tergites T2 to T4 yellowish with very distinct black stripes or entirely dark brown, T5 with broader stripe or completely black, T6 shiny black ( Fig. 4 ); cercal ventral lobe (secondary clasper) fused to cerci with two or three very large curved black medial teeth on the internal margin; surstylus quadrate with a lateral row of strong, short prensisetae on the outer and inner margins (with two medial prensisetae in D. chocolata sp. nov. ), the innermost one significantly long; hypandrial median process tapering; aedeagus hirsute with fine cuticular scales; posterior parameres as long as aedeagus (slightly shorter in D. chocolata sp. nov. ), with finely serrated margins ( Figs 3A , 5 ). Female abdominal tergites lighter than male’s or concolorously dark ( Fig. 4 ). Remarks The original ‘ D. seguyi species complex’ was renamed ‘ D. bakoue species complex’ without new definition or justification ( Rafael 1984 ). We consider here the ‘ D. bakoue species complex’ to be a junior synonym for the ‘ D. seguyi species complex’. Chassagnard et al. (1997) discussed a ‘ D. seguyi species complex’ comprising D. curta , D. seguyi and D. seguyiana in a table legend, but considered all these species members of the ‘ D. bakoue species complex’ in the main text. We restrict here the ‘ D. seguyi species complex’ to four monophyletic species, while placing D. seguyiana in the ‘ D. tsacasi species complex’ (see below). David et al. (2014) referred to three of these species as geographical strains of D. malagassya with partial reproductive isolation. Closer examination of their material revealed, however, that each strain was a distinct species (namely, D. seguyi , D. malagassya and D. curta ). Taxon content D. seguyi Smart, 1945 . D. malagassya Tsacas & Rafael, 1982 . D. curta Chassagnard & Tsacas, 1997 . D. chocolata sp. nov.