Taxonomy of Serpulidae (Annelida, Polychaeta): The state of affairs
Author
Ten Hove, Harry A.
Author
Kupriyanova, Elena K.
text
Zootaxa
2009
2009-03-16
2036
1
126
journal article
11755334
15888B41-A000-4611-BEC8-F9359D1149CD
40.
Semivermilia
ten
Hove, 1975
(
Fig. 45
)
Type-species
:
Vermiliopsis pomatostegoides
Zibrowius, 1969b
Number of species: 8
Tube triangular to sub-triangular in cross-section, keels present, otherwise variable; without flaring peristomes; granular overlay absent. Operculum inverse conical with chitinous endplate, more often a cap or series of diabolo-like plates; sometimes with terminal spine. Peduncle inserted as second radiole, cylindrical in cross-section; constriction present. Pseudoperculum absent. Radiolar arrangement short pectinately, up to 7 radioles per lobe. Inter-radiolar membrane absent. Branchial eyes may be present. Stylodes absent. Mouth palps present. (5-)7 thoracic chaetigerous segments. Collar tri- to penta-lobed, tonguelets absent. Thoracic membranes end at chaetiger 2. Collar chaetae limbate.
Apomatus
chaetae present in posterior thoracic segments (
Fig. 45A
). Thoracic uncini saw- to-rasp-shaped; with about 15 teeth in profile view, 1 tooth at the apex of the uncinus to 5 teeth in the row above the wide gouged peg (dental formula e.g., P:5:3:2:2:1:1:1:1:1:1:1:1:1:1;
Fig. 45B
). Triangular depression absent. Abdominal chaetae flat narrow geniculate, with rounded teeth on edge (
Fig. 45D, E
); abdominal uncini smaller than thoracic ones, entirely rasp-shaped, with about 13 teeth in profile view, up to 8 teeth in a row (
Fig. 45C
). Achaetous anterior abdominal zone, if present, very short. Long posterior capillary chaetae absent. Posterior glandular pad may be present.
Remarks.
Within the group of
Vermiliopsis
-like genera,
Zibrowius (1972b
,
1973a
, b) distinguished 4 genera (
Bathyvermilia
,
Metavermilia
,
Pseudovermilia
, and
Vermiliopsis
) and a group of aberrant species that he termed?
Vermiliopsis
. Ten
Hove (1975)
erected a new genus
Semivermilia
for part of the latter (leaving the attribution of?
Vermiliopsis glacialis
Monro, 1939
,
V. notialis
Monro, 1930
, and
V. eliasoni
Zibrowius, 1970a
uncertain) and provided a table that allows distinguishing between the five genera above and
Neovermilia
.
In small specimens/species, such as
Semivermilia pomatostegoides
(this paper), and certainly
S. cribrata
(as
Josephella carenata
Zibrowius, 1968a: 176
) the number of thoracic chaetigers may be 5 or 6. Contrary to statements in the literature that the peduncle is the first radiole (e.g.,
Zibrowius (1968a)
for
S. crenata
as
Vermiliopsis undulata
, respectively
Zibrowius (1969b)
as
V. pomatostegoides
), we checked material (
Canary Islands
, SW coast of La Palma, Punta del Hombre,
8–12 m
,
May 29, 1980
, CANCAP Sta. 4.D10, ZMA V.Pol. 4213), and found it to be the second normal radiole, or to be inserted just below the first and second normal radiole (thus probably migrated down from the second position). The character is difficult to observe, and may be subject to prejudice.
FIGURE 45.
SEM micrographs of chaetae in
Semivermilia elliptica
. Japan, Shirohama, Seto Marine Biological Laboratory, marine aquarium, legit E. Kupriyanova, det. H.A. ten Hove, SAM E3664. A—4
th
thoracic bundle, B—uncini of 4
th
thoracic chaetiger, C—abdominal uncini, D—anterior abdominal chaeta, E—posterior abdominal chaetae.
1.
Semivermilia agglutinata
(
Marenzeller, 1893
)
, Benghazi, Mediterranean; bathyal
2.
Semivermilia crenata
(O.G.
Costa
, 1861
)
, Mediterranean, Mauretania
3.
Semivermilia cribrata
(O.G.
Costa
, 1861
)
, Mediterranean, Mauretania
4.
Semivermilia elliptica
Imajima, 1978
, Izu Island,
Japan
5.
Semivermilia parapomatostega
Wu & Chen, 1981a
, South
China
Sea
6.
Semivermilia pomatostegoides
(
Zibrowius, 1969b
)
, Tripolis, Mediterranean, Central Atlantic, Indo-West Pacific
7.
Semivermilia torulosa
(delle Chiaje, 1822), Mediterranean, Mauretania
8.
Semivermilia uchidai
Imajima & ten Hove, 1986
,
Solomon Islands
,?
Seychelles
.