A monograph of the genus Maladera Mulsant & Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) Author Fabrizi, Silvia 0000-0003-3524-7153 ahrens.dirk_col@gmx.de Author Liu, Wan-Gang 0000-0003-4788-7967 liuwangang@ieecas.cn Author Bai, Ming 0000-0003-3676-6828 liuwangang@ieecas.cn Author Yang, Xing-Ke 0000-0003-3676-6828 liuwangang@ieecas.cn Author Ahrens, Dirk 0000-0003-3524-7153 ahrens.dirk_col@gmx.de text Zootaxa 2021 2021-02-03 4922 1 1 400 journal article 8257 10.11646/zootaxa.4922.1.1 188411de-569c-4408-a016-1a04b858c142 1175-5326 4496316 D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C Maladera perniciosa ( Brenske, 1898 ) Figures 14 A–E, 93 Autoserica perniciosa Brenske, 1898: 336 . Maladera ( Cephaloserica ) perniciosa : Ahrens 2004a: 276 ; Krajcik 2012: 155 ; Ahrens & Fabrizi 2016: 242 . Type material examined. See Ahrens (2004a) . Additional material examined. China : 1 ♂ “ Pantiange , Weixi , Yunnan , 29.VII.1981 , 2500m , leg. Wang Shuyong” ( IZAS ) , 2 ♂♂ “ Huoditang , Ningshan , Shaanxi , 30.VI.1999 , 1580m, leg. Yuan Decheng ” ( IZAS ) , 2 ex. “China N Sichuan Jiuzhaigou 2000 m 16. Jun 1992 M. Bok lgt.” ( SMNS ) , 1 ex. “ China : Yunnan Province , Tengchong , 3.-6.VI.2007 near long distance bus station, 25°00.3’N 098°29.3’E , 1625m , J. Hajek & J. Ruzicka leg.” ( NMPC ) , 1 ex. “ China : Yunnan Province , Shanzhi env., 22.-24.VI.2007 Jizu Shan Mt. , 25°57.7’N 100°23.6’E , 2180m , J. Hajek & J. Ruzicka leg.” ( NMPC ) , 5 ex. “ China : Yunnan Province , Dali old city, 31.V.-2.VI.2007 , 25°41.4’N 100°09.8’E , 2030m , J. Hajek & J. Ruzicka leg.” ( NMPC ) , 106 ex. “ Tibet : Ma-ma-luo, Zayu county , 15-26 June 2010 local collector of Li Jingke” ( ZFMK ) , 31 ex. “ China Yunnan 2000-3000m 27.20N 100.11E Habashan mt. SE slope 10.-13.7. David Kral leg. 92/ coll. Milan Nikodym , Praha ” ( ZFMK , NMPC ) , 2 ex. “ China , N-Yunnan 27°18’N 100°13’E Jinsha r. vall. 1900M Daju, Hutiao gorge lgt. D. Kral 15-17/7’90” ( ZFMK ) , 2 ex. “ China Yunnan 2000-3000m 27.20N 100.11E Habashan mts. SE slope 10-13/7. Vit Kuban leg. 92” ( ZFMK ) , 6 ex. “ China ( Yunnan ) Dali Bai Aut. Pref. , foothills of Diacang Shan W of Dali 2050m 25°42‘12‘‘N / 100°08‘26‘‘E (stone and gravel pit) 12.VI.2007 D.W. Wrase [34]” ( ZFMK ) , 12 ex. “ China : Sichuan ; Wolong Reserve , Sigulian Shan , 31º09‘N 103º06‘E v.2006 , 1500- 1800m V. Siniaev ” ( ZFMK ) , 1 ♂ “ China Yunnan 27.-28.VI. Daju , 50 km M Lijiang 27°21’N 100°19’E E. Jendek leg. 1992” ( NHMW ) , 1 ♂ “ China : E-Yunnan; Damaidi 2500m , Guangnan near Vietnam VII-2003 leg. Li et al. ” ( ZFMK ) . Vietnam : 1 ex. “ Tonkin Montes Mauson April , Mai 2-3000‘ H. Fruhstorfer ” ( ZMHB ) , 2 ex. “N-Vietnam Cao Bang Prov. , vic. Tinh Tuc , Nui Pia Oac Nat. Res. 13.V.2013 , 22°36‘50‘‘N , 105°36‘50‘‘E 14-1800m leg. A. Skale ” ( CASH ) , 2 ex. “N-Vietnam Vinh Phuc Prov. vic. Tam Dao Town , Tam Dao NP, 02.-05.V.2013 , 21°27‘N , 105°38‘E 700-1000m , A. Skale ” ( CASH ) , 1 ex. “N-Vietnam Cao Bang Prov. , vic. Vin Den , Nui Pia Oac Nat. Res. 06.-10.V.2013 , 22°33‘53‘‘N , 105°52‘53‘‘E 900-1300m A. Skale ” ( CASH ) , 2 ex. “N-Vietnam Cao Bang Prov. , vic. Vin Den , Nui Pia Oac Nat. Res. 10.-15.V.2013 , 22°33‘53‘‘N , 105°52‘53‘‘E 900-1300m A. Skale ” ( CASH ) , 1 ♂ “Vietnam-N Tuyen Quang pr. SE-E env. Of Na Hang , 22°17’30’’N 105°26-28’E , 200-700m L. Dembicky leg., 1.- 12.V.2010 ” ( ZFMK ) . Thailand : 1 ex. “Thai-N, Nan Prov. , Bo Klua , 700 m , 19°8’N 101°10’E , 28.iv.-1.v.1999 , D. Hauck leg.” ( CP ) , 2 ex. “N-Thailand 17.VII.1990 Doi Inthanon, leg. Malicky ” ( ZSM ) , 4 ex. “N-Thailand V.1990 Doi Inthanon, leg. Malicky ” ( ZSM ) , 1 ♂ “ Thailand , Mae Hong Son prov. , Kiwlom pass near Soppong , WGS 84; 19°26’N 98°19’E , 23.vi.-2.vii.2002 , alt. 1400m + 50m , lgt. Fouque R.+ H. Kozich , J.” ( NMPC ) , 1 ♂ “N. Thailand : Angkhai village , Samoeng Dist. Chiang Mai Prov. , 9.-11.v.1999 K. Masumoto leg.” ( ZFMK ) . Laos : 1 ex. “ Laos : Sayaboury Prov. Sayaboury 5.V.1966 / Native Collector Bishop Museum / Autoserica rubescens Mos. det. G. Frey 1971” ( CF ) , 2 ex. “NE-Laos: Hua Phan prov. , Ban Saleui , Phou Pan ( Mt. ) - 20°12’N , 104°01’E ; 14.iv.-15.v.2012 ; 1300-1900m ; leg. C. Holzschuh Ankauf ZFMK Bonn 2012/13” ( ZFMK ) , 2 ex. “Laos-NE Hua Phan prov. , 20°12’N , 104°01’E , Phu Phan Mt. , 1500-1900m , 17.5.- 3.6.2007 , leg. Vit Kuban ” ( ZFMK ) , 3 ex. “NE-Laos: Hua Phan prov. , Ban Saleui , Phou Pan ( Mt. ) - 20°12’N , 104°01’E ; 11.iv.-15.v.2012 ; 1300-1900m ; leg. C. Holzschuh Ankauf ZFMK Bonn 2012” ( ZFMK ) , 1 ex. “Laos-NE, Houa Phan prov., 20°13’09- 19’’N 103°59’54’’- 104°00’03’’E , 1480-1510m Phou Pane Mt., 22.IV.-14.V.2008 Vit Kuban leg. ( NMPC , ZFMK ), 1 ♂ “X-DA4702 Laos Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) 20°12’N , 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Maladera perniciosa” ( ZFMK ), 1 ♂ “X-DA4918 Laos Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) 20°12’N , 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Maladera perniciosa ” ( ZFMK ), 1 ♂ “X-DA4923 Laos Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) 20°12’N , 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Maladera perniciosa ” ( ZFMK ), 1 ♂ “X-DA4699 Laos Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) 20°12’N , 104°01 ’ 15.- 30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Maladera perniciosa ” ( ZFMK ), 1 ♂ “X-DA4830 Laos Hua Phan prov. ; Ban Saleui , Phou Pan ( Mt. ) 20°12’N , 104°01 ’ 15.-30.iv.2014 leg. C. Holzschuh—ZFMK Ankauf 2014 Maladera perniciosa ” ( ZFMK ) . FIGURE 14. A–E: Maladera perniciosa ( Brenske, 1898 ) (Vietnam: Lao Cai: Sa Pa); F–J: M. sontica ( Brenske, 1898 ) (Laos: Louangnamtha: Namtha); K–O: M. verticalis ( Fairmaire, 1888 b) (China: Shaanxi: Houzhenzi). A, F, K: aedeagus, left side lateral view; D, I, N: aedeagus, right side lateral view; B, G, L: aedeagus, dorsal view; C, H, M: left parameres, ventral view; E, J, O: habitus. Scale: 0.5 mm. Habitus not to scale. Remarks. The species was exhaustively treated (including redescription) by Ahrens (2004a , p. 276). Distribution. See map ( Fig. 93 ) and Table 1 .