A taxonomic revision of two genera, Pleurolobus and Sohmaea (Leguminosae) in Thailand and Indo-China
Author
Saisorn, Witsanu
School of Science, Walailak University, Nakhon Si Thammarat 80161, Thailand.
Author
Chantaranothai, Pranom
Department of Biology & Centre of Excellence on Biodiversity (BDC), Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand.
text
Phytotaxa
2022
2022-11-23
573
2
231
246
http://dx.doi.org/10.11646/phytotaxa.573.2.4
journal article
194617
10.11646/phytotaxa.573.2.4
80d52cdf-fbd6-4aa4-b9f1-c6e8ac9dc473
1179-3163
7349951
4.
Sohmaea teres
(Wallich 1831: 196 ex
Bentham 1852: 225
) H.Ohashi & K.Ohashi (2018b: 163)
.
Fig. 2C–D
.
≡
Desmodium teres
Wallich ex Bentham
Type:—
MYANMAR
.
Taong Dong
,
Wallich
5694
(
lectotype
K-W!
K001121790
, designated by
Rafidah
et al.
(2020)
, isolectotypes BM! BM000946774, CAL,
n.v.
, G!, K! K000858870, K-W! K001121791).
≡
Meibomia teres
(Wall. ex Benth.)
Kuntze (1891: 198)
.
=
Desmodium zonatum
Meeuwen (1961: 97)
,
non
Miq. (1855: 250). Specimen cited:—
Dorgeh 1755
(L, n.v.),
fide
Ohashi (2004)
.
Herb, up to
2 m
high; stems and twigs terete, sparsely to densely uncinate hairy.
Leaves
unifoliolate, alternately arranged; stipules narrowly triangular, 5–12 ×
2–3 mm
, apex long acuminate to caudate, surface glabrescent; petioles
0.3–0.8 cm
long, ascending pubescent and minutely uncinate.
Leaflets
: stipels narrowly triangular, 2–5 ×
0.5–1 mm
, apex long acuminate, surface glabrescent, margin not fimbriate; petiolules
1.5–3 mm
, ascending pubescent and uncinate hairy; leaf blade lanceolate to narrowly lanceolate or oblong, (3–)9–20 × (0.5–)
2.5–7 cm
, apex attenuate, base obtuse, margin entire, upper surface nearly glabrous or sparsely uncinate hairy, with straight hairs on mid- and lateral veins, lower surface spreading uncinate hairy, with straight hairs on mid- and lateral veins; lateral veins 9–14 per side.
Inflorescences
10–40 cm
long, terminal or axillary; rachis densely uncinate hairy.
Primary bract
narrowly ovate, 3.5–4 ×
1–1.5 mm
, apex long acuminate to attenuate, surface pubescent, conspicuously veined, enclosing 5-immature flowers and secondary bract.
Secondary bract
narrowly ovate to narrowly triangular, 1.5–2 ×
0.5 mm
, pubescent.
Flowers
6–7 mm
long, 5-flowered fascicles; bracteoles absent; pedicels
4–5 mm
long, densely uncinate and straight hairy (on lower half part).
Calyx
yellowish green, ca.
4 mm
long, base cuneate; outside ascending pubescent and minutely uncinate, inside glabrescent, tube ca.
1.5 mm
long; teeth
2–3 mm
long, acuminate to needle-like at the apex, prominently longer than tube length, two lateral teeth shorter than others, lower tooth longest, upper one entire.
Corolla
standard white, wings and keels violet to blue; standard broadly obovate to oblanceolate, 4.5–8 ×
3.5–5.2 mm
, apex obtuse to rounded, base attenuate, not auriculate, claw
0.3–1 mm
long; wings slightly oblong, 3.5–6 ×
1–2 mm
, apex obtuse, base auriculate, ca. 0.3 ×
0.5 mm
, claw
0.5–1 mm
long; keels slightly curved, 4–7 ×
1–1.5 mm
, prominently longer than wings, apex acute to slightly obtuse, base shortly auriculate, claw ca.
2.5 mm
long.
Stamens
4–7.5 mm
long; anthers ellipsoid, ca. 0.3 ×
0.2 mm
.
Gynoecium
4.5–9 mm
long; ovary oblong, laterally compressed, pubescent, stipitate,
0.5–1 mm
long, 10–13-ovulate; style
1.5–3 mm
long, pubescent; stigma minutely capitate.
Pods
light brown, indehiscent, 8–11-articulate, subterete,
4–6.5 cm
long,
1–1.5 mm
wide, surface densely uncinate, not reticulate-patterned (but longitudinal striate), both sutures less constricted between articles and thickened; isthmus ca. 4/5 as broad as the pod; articles narrowly oblong to linear,
5–6 mm
long; pod stipe
1.5–2 mm
long, glabrescent; fruiting pedicels
5–7 mm
long, ascending pubescent and uncinate hairy.
Seeds
brown, transversely oblong, 4.5–5 ×
1 mm
, ca.
0.7 mm
thick.
Distribution
:—
Myanmar
,
Thailand
,
Malaysia
, and
Indonesia
.
Ecology
:—In sunny places of limestone mountain and evergreen, dry evergreen, and mixed deciduous forests;
50–700 m
elev.
Phenology
:—Flowering in September–November. Fruiting in October–January.
Notes
:—
Sohmaea teres
is a new record species for
Cambodia
.
Specimens examined
:—
THAILAND
.
Chiang Mai
:
Mae Rim
, QBG,
BGO.
Staff
4828
(
QBG-2
sheets),
Doi Chiang Dao
,
480 m
,
21 Oct. 1955
,
Garrett
1460
(
K
,
L
, P-2 sheets),
Doi Chiang Dao
,
600–1300 m
,
25 Sept. 1971
,
Murata
et al. T-14911
(
KYO
)
,
Doi Chiang Dao
,
600–1300 m
,
25 Sept. 1971
,
Murata
et al. T-14936
(
KYO
)
,
Chiang Dao
,
Ban Bing Kong
,
500 m
,
29 Oct. 1987
,
Maxwell
87-1304
(
CMU
,
L
)
,
Doi Suthep
,
630 m
,
17 Oct. 1999
,
Srapratet
24
(
CMUB
)
,
Fang
,
Doi Pha Hom Pok National Park
,
21 Dec. 2014
,
Saisorn
352
(
KKU
)
,
Mae Rim
, QSBG,
860 m
,
13 Nov. 2014
,
Tanming
732
(
QBG
)
&
Chiang Dao
,
Tham Chiang Dao
,
350 m
,
15 Feb. 1958
,
SØrensen et al. 1137
(
C
)
;
Chiang Rai
:
Doi Pha Hom Pok
,
700 m
,
15 Sept. 1967
,
Iwatsuki
et al. T-9724
(
KYO
)
&
Pan
,
Doi Luang National Park
,
610 m
,
3 Nov. 2015
,
Muangyen
214
(
QBG
)
;
Phayao
:
Mae Chai
,
Doi Luang National Park
,
800 m
,
4 Nov. 2015
,
Muangyen
318
(
QBG
)
,
Doi Luang National Park
,
600 m
,
4 Nov. 2015
,
Muangyen
327
(
QBG
)
&
Mueang
Phayao
,
Doi Luang National Park
,
575–625 m
,
25 Sept. 1997
,
Petrmitr
123
(
BKF
,
CMUB
)
;
Nan
:
Tham Pha Tok
,
250 m
,
16 Nov. 1993
,
Larsen
et al. 44405
(
AAU
)
;
Lampang
:
Chae Hom
,
325 m
,
26 Oct. 2012
,
Maxwell
12-279
(
CMUB
,
QBG
)
, en route from Pang La to Huai
Tak
,
350 m
,
24 Sept. 1967
,
Shimizu
et al. T-10752
(
BKF
,
KYO
)
&
without locality,
250 m
,
22 Nov. 1922
,
Winit
811
(
ABD
,
BK
,
K
)
;
Sukhothai
:
Si Satchanalai
,
Ban Mae San
,
17 Nov. 2015
,
Maknoi
8391
(
QBG
)
&
Ban Mae San
,
28 Jan. 2016
,
Maknoi
8514
(
QBG
)
;
Nakhon Sawan
:
Ban Na Kluay
,
50 m
,
7 Nov. 1924
,
Kerr
4537
(
ABD
,
BK
,
BM
,
E
,
K
)
;
Phetchabun
:
Nam Nao National Park
,
3 Nov. 2013
,
Mattapha
448
(
KKU
)
&
Nam Nao National Park
,
18 Oct. 2004
,
Mattapha
696
(KKU-2 sheets)
;
Loei
:
Pha Nok Khao
,
250–350 m
,
6 Dec. 1970
,
Charoenphol
et al. 4564
(
AAU
,
BKF
,
C
)
&
Wat Tham Pha Sawan
,
381 m
,
14 Nov. 2011
,
Clark
et al. 240
(
K
,
QBG
)
;
Khon Kaen
:
Phu Pha Man National Park
,
25 Oct. 2007
,
Norsaengsri
&
Lakoet
2807
(
QBG
)
,
Phu Pha Man National Park
,
410.6 m
,
22 Nov. 2007
,
Norsaengsri
&
Lakoet
3069
(
QBG
)
,
Phu Pha Man National Park
,
467 m
,
24 Dec. 2011
,
Norsaengsri
&
Tathana
8403 (
QBG
)
,
Wiang Kao
,
Phu Wiang National Park
,
11 Nov. 2014
,
Saisorn
et al. 294
(
KKU
)
,
Wiang Kao
,
Phu Wiang National Park
,
15 Feb. 2015
,
Saisorn
376
(
KKU
)
,
Chum Phae
,
Dong Lan
,
300 m
,
22 Nov. 1958
,
Smitinand
4919
(
BKF
,
L
)
&
Chum Phae
,
Dong Lan
,
400 m
,
22 Dec. 1958
,
SØrensen et al.
6122 (
C
)
;
Nakhon Ratchasima
:
Pak Chong
,
600–650 m
,
20 Nov. 1982
,
Koyama
et al. T-30147
(
KYO
)
,
Pak Chong
,
700–730 m
,
Koyama
et al. T-30094
(
KYO
)
&
without locality,
18 Dec. 1962
,
Phengklai
584
(
C
,
K
,
L
)
;
Kanchanaburi
:
Wang Po
,
26 Oct. 1969
,
Kasem
648
(
BK
,
L
,
SING
)
,
Srisawat
,
580–600 m
,
26 Nov. 1982
,
Koyama
et al. T-30304
(
KYO
)
,
Sai Yok
,
Mahidol University
Kanchanaburi
Campus
,
225 m
,
20 Dec. 2005
,
Maxwell
05-739
(
CMUB
)
,
Si Sawat
,
Ban Khao Lek
,
500–600 m
,
21 Oct. 1994
,
Nisit
94102118
(
BK
)
,
Si Sawat
,
Ban Khao Kaeng Khaep
,
500 m
,
2 Nov. 1979
,
Shimizu
et al. T-21399
(
BKF
,
KYO
,
L
)
,
Si Sawat
,
Erawan National Park
,
100–300 m
,
3 Nov. 1979
,
Shimizu
et al. T-21540
(
BKF
,
KYO
)
,
Si Sawat
,
Erawan National Park
,
100–300 m
,
3 Nov. 1979
,
Shimizu
et al. T-21567
(
KYO
)
,
Sai Yok
,
160 m
,
6 Nov. 1979
,
Shimizu
et al. T-21760
(
BKF
,
KYO
)
,
Sai Yok
,
Near Sai Yok Arboretum
,
160 m
,
6 Nov. 1979
,
Shimizu
et al. T-21829
(
KYO
)
&
Si Sawat
,
150 m
,
5 Nov. 1971
,
van Beusekom
et al. 3413
(
BKF
,
C
,
K
, KYO, L-2sheets,
P
)
;
Prachuap Khiri Khan
:
Kui Buri
,
Sam Roi Yot
,
23 Oct. 1973
,
Sutheesorn
2703
(
BK
)
;
Lop Buri
:
Chai Badan
,
280 m
,
4 Dec. 1982
,
Koyama
&
Terao
T-30595
(
BKF
,
KYO
)
&
Subchampa Hill
,
170–190 m
,
19 Nov. 1984
,
Murata
et al. T-50933
(
BKF
,
KYO
)
;
Saraburi
:
Muek Lek
,
200 m
,
10 Nov. 1924
,
Marcan
1867
(
ABD
,
BK
,
BM
,
KKU
)
;
Chon Buri
:
Si Racha
,
Khao Khiao
,
300 m
,
18 Dec. 1974
,
Maxwell
74-1077
(
BK
)
&
Si Racha
,
Khao Khiao
,
100 m
,
20 Oct. 1975
,
Maxwell
75-1052
(
AAU
,
BK
,
L
)
;
Chanthaburi
:
Pong Nam Ron
,
700 m
,
11 Oct. 1955
,
Sangkhachand
528
(
BKF
,
L
)
,
Pong Nam Ron
,
Ban Thalee
,
200 m
,
24 Oct. 1956
,
Smitinand
3616
(
BKF
)
&
without locality,
200 m
,
28 Nov. 1956
,
Smitinand
3648
(
BKF
,
L
)
;
LAOS
.
Luang Prabang
:
Massie
s.n
(
Dy Phone
et al.
, 1994);
CAMBODIA
.
Mondulkiri
:
Sen Monorom
,
527 m
,
5 Nov. 2016
,
Yahara
et al. 7382
(
FU
)
;
VIETNAM
.
Annam
:
Ca-na
,
Phanrang
,
500 m
,
22 Nov. 1923
,
Poilane
8652
(P-2 sheets)
.