Revisions to the Andrena fauna of north-western Africa with a focus on Morocco (Hymenoptera: Andrenidae)
Author
Wood, Thomas James
text
European Journal of Taxonomy
2023
2023-12-21
916
1
1
85
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2381/10463
journal article
10.5852/ejt.2023.916.2381
2118-9773
10453460
0DC587F6-9DAA-4F6E-BA2A-AD528990BA24
Andrena
(
Chrysandrena
)
testaceipes
Saunders, 1908
Fig. 12
Andrena
(
Chrysandrena
)
testaceipes
Saunders, 1908: 196
,
♀
(
Algeria
: NHMUK).
Andrena
(
Chrysandrena
)
colorata
Alfken, 1929: 269
,
♀
♂
(
Libya
: ZMHB).
Andrena
(
Chrysandrena
)
rubricorpora
Wood, 2021: 465
,
♀
(
Tunisia
: OÖLM).
Syn. nov.
Andrena cirtana
– auctorum, nec.
Lucas, 1849
.
Material examined
ALGERIA
•
♀
,
lectotype
of
A. testaceipes
(
Fig. 12A–D
);
Constantine
;
20 May 1895
;
A.E. Eaton
leg.;
NHMUK
.
LIBYA
•
1 ♀
;
Wadi Kuf
;
1 Apr. 1954
;
K.M. Guichard
leg.;
NHMUK
.
MOROCCO
•
5 ♂♂
,
1 ♀
;
Fès-Meknès
,
Ahermoumou, P
5407, immediately
NW of Kassioua
;
900 m
a.s.l.
;
15 May 2022
;
T.J. Wood
leg.;
TJWC
•
1 ♀
; SE of
Asni
,
Oukaimeden
;
2600 m
a.s.l.
;
24 Jul. 1985
;
K.M. Guichard
leg.;
NHMUK
.
TUNISIA
•
♀
,
holotype
of
A. rubricorpora
;
Zaghodan Mts
;
14 May 1993
;
J. Batelka
leg.;
OÖLM
•
1 ♀
,
paratype
of
A. rubricorpora
; same collection data as for holotype;
OÖLM
.
Remarks
Confusion has surrounded this taxon since
Warncke (1967)
used
A. cirtana
Lucas, 1849
to apply to what he considered to be a red-marked species of
Margandrena
(see
Warncke 1974
for subgeneric affiliation).
Grünwaldt (1976)
noted that the name
A. cirtana
cannot apply to this taxon, as the
holotype
is a male, and the male of
A. cirtana
was described as having a yellow-marked clypeus, whereas Warncke’s taxon has the clypeus black. Grünwaldt argued that
A. testaceipes
was therefore the priority name.
Warncke (1967)
designated a
lectotype
for
A. testaceipes
(
Fig. 12A–D
) which fits Saunders’ description. Importantly, this taxon does not match the criteria for the subgenus
Margandrena
, and instead falls in
Chrysandrena
due to the lack of a strong pronotal angle, and the tibial scopa is composed of strongly plumose hairs. The genital capsule of
A. testaceipes
was illustrated by
Scheuchl (2010)
, where the typical
Euandrena
/
Chrysandrena
form can
be seen. This unclear phylogenetic position led to the description of
A.
(
Chrysandrena
)
rubricorpora
Wood, 2021
from
Tunisia
(Wood 2021), as it was not clear that
A. testaceipes
was actually a member of
Chrysandrena
. Genetic samples from
Morocco
clearly place
A. testaceipes
within
Chrysandrena
; it is therefore transferred there, and
A. rubricorpora
is synonymised with it.
Fig. 12.
Andrena
(
Chrysandrena
)
testaceipes
Saunders, 1908
, lectotype, ♀ (NHMUK).
A
. Label details.
B
. Lateral habitus.
C
. Face, frontal view.
D
. Terga, dorsal view.
One issue remains, which is the status of the true
A. cirtana
. At the moment, its identity is unclear, as
Gusenleitner & Schwarz (2002)
only listed this taxon in its sensu auctorum. Location and examination of the
type
series in the MNHN collection is necessary, as it potentially has priority over an existing name given its early description date of 1849. No action will be taken until the
type
series is located.
Distribution
Morocco
,
Algeria
,
Tunisia
,
Libya
,
Egypt
(
Gusenleitner & Schwarz 2002
).