Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr Author Spiegel, Didier Van Den Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be text Zoosystema 2007 29 1 7 49 journal article 10.5281/zenodo.5392079 1638-9387 5392079 Glyphiulus pergranulatus n. sp. ( Figs 21-23 ) TYPE MATERIAL. — China . Guizhou Prov. , Guanling County , Huajiang , Da Dong Cave , 4.IV.2004 , leg. L. Latella & Ghen Hu , holotype ♂ ( MNHN GA 041 ) ; paratypes 1 ♂ ( MNHN GA 041 ) ; 1 ♀ ( SEM ). — Guizhou Prov. , Guanling County , Huajiang , An Jia Da Dong Cave , 4.IV.2004 , leg. L. Latella & R. Zorzin , paratype 1 ♀ ( MNHN GA 041 ) . ETYMOLOGY. — To emphasize the close affinities to G. granulatus . DIAGNOSIS. — Characterized by the somewhat larger ocellaria, the underdeveloped tergal crests (becoming almost obliterated in the median part of the collum), coupled with the medium body size, slightly elongated legs, simple anterior gonopods, etc. DESCRIPTION FIG. 23. — Glyphiulus pergranulatus n. sp. , ♂ paratype: A , antenna; B , legs 1, caudal view; C , legs 2, caudal view; D , leg 3; E , midbody leg; F , anterior gonopods, caudal view; G , posterior gonopod, subfrontal view; H , same, frontal view. Scale bar: 0.2 mm. Length of holotype and of paratype ♀ from An Jia Da Dong about 30 mm , of paratype ♂ about 33 mm , width 1.5, 1.5 and 1.7 mm , with 56p+2a+T, 67p+1a+T and 57p+2a+T, respectively. Coloration, antennae ( Figs 21A ; 23A ), segment shape ( Fig. 22D ), carinotaxy patterns ( Fig. 21 ), male legs 1-3 ( Fig. 23 B-D), etc., as in G. granulatus , but tergal crests somewhat underdeveloped ( Figs 21 ; 22B, D ), almost obliterated in median part of collum ( Fig. 21A, B ). Ocellaria rounded-triangular, black, each with 23-25 poorly convex ocelli. Claws slightly longer, spine at base slightly shorter ( Fig. 22C ); legs about 4/5 as long as midbody height ( Fig. 23E ). Epiproct with a faint but evident middorsal tubercle ( Fig. 22B ); hypoproct more narrowly emarginate at caudal margin ( Fig. 22A ). Anterior gonopod coxosternum particularly simple ( Fig. 23F ). Posterior gonopods with a typical flagelliform and plumose distal process ( Fig. 23G, H ). REMARKS Even though taken from caves, this species seems to represent only a troglophile at most. However, this cave harbours another, apparently troglobitic congener from a different species group, to be described elsewhere.