Leptoomidae, a new family of Eocene fossil Chalcidoidea (Hymenoptera), and family classification of Eocene fossil genera originally described in Neanastatinae (Eupelmidae)
Author
Gibson, Gary A. P.
Honorary Research Associate, Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, K. W. Neatby Bldg., 960 Carling Avenue, Ottawa, Ontario, CANADA, K 1 A 0 C 6.
Author
Fusu, Lucian
Al. I. Cuza University, Faculty of Biology, Bd. Carol I, Nr 11, 700506 Iasi, ROMANIA.
text
Zootaxa
2023
2023-07-19
5318
2
195
216
http://dx.doi.org/10.11646/zootaxa.5318.2.2
journal article
10.11646/zootaxa.5318.2.2
1175-5326
8162469
CF8E0B91-9AF7-4075-963D-9BC977B41852
Neanaperiallus
Gibson
(
Figs 1A
,
2B
,
4
,
5
)
Neanaperiallus
Gibson, 2009: 200‒203
.
Included species
.
Neanaperiallus janzeni
Gibson
and
Neanaperiallus defunctus
Fusu
sp. nov.
Diagnosis
. Head and mesosoma non-metallic, brown. Head with inner margins of eyes apparently not divergent ventrally or at least not conspicuously divergent (not visible in
N. janzeni
and direct frontal view not possible for
N. defunctus
,
Fig. 5E
); clypeal area concave and labrum perpendicular to clypeus (
i. e.
not visible in direct frontal view of clypeus,
cf
.
Fig. 5E
and inset in 5E); occipital carina
∩
-shaped (
Fig. 5F
;
Gibson 2009
, fig. 59). Mandible not visible in
N. janzeni
but apparently tridentate in
N. defunctus
. Antenna with lower margin of torulus in line with or slightly ventral to lower margin of eyes; 13-segmented (1:1:8:3) with 8 slightly transverse to slightly elongate funiculars and with mps on fu
3
‒fu
8
(
Figs 4B, C, F
;
Gibson 2009
, fig. 58); clava elongate-ovoid, more than twice as long as wide, and apical clavomere with minute micropilose sensory region apically, appearing as rounded terminal button (
Fig. 4C
: tb). Mesoscutellar-axillar complex with axillae equilateral to somewhat elongate-triangular, with their inner angles slightly separated and mesoscutellum truncate basally when mesonotum not flexed (
Figs 4G, H
) (flexed condition unknown); mesoscutellum with sculpture along apical margin aligned, differentiating a thin carina (
Figs 4E, 4H
,
5H
: smc) separating lunate, down-sloped region apically. Metanotum with dorsellum an oblique band projecting over apex of mesoscutellum (
Fig. 5H
) but the two sclerites separated by a narrow gap (
Fig. 5H
: double arrow in inset). Prepectus flat (
Fig. 4F
: pre). Acropleuron enlarged posteriorly to metapleuron and posteroventrally to dorsolateral level of mesocoxa, with crenulate acropleural sulcus directed obliquely toward posterodorsal angle of prepectus over most of length (
Fig. 1A
) or only over about anterior one-third (
Fig. 5A
) but differentiating acropleuron from comparatively large mesepisternal region ventral to acropleural suture (
Figs 1A
,
5A
). Mesopectus in ventral view with posterior margin abutting base of mesocoxa (
Fig. 2B
) though mesocoxa with triangular basolateral cavity opposite angle formed between acropleuron and mesopectus (
Fig. 2B
;
Gibson 2009
, fig. 61). Fore wing disc with bare band beyond basal fold separated from parastigma by region of setae (
Figs 5G, H
;
Gibson 2009
, fig. 62), parastigma not separated from marginal vein by hyaline break though apparently slightly less pigmented distally (
Figs 5G, H
). Protibia apparently without dorsal spicules but with dorsoapical spicules (
Fig. 5A
, inset); mesotibia with spine-like pegs apically (
Fig. 5C
).
Remarks
. The extended generic diagnosis given above for
Neanaperiallus
is to supplement the description in
Gibson (2009)
after the discovery of a second fossil that represents a second species in the genus, and to have a comparative description with
Leptoomus
. As noted above, the state of preservation and the inability to examine the two available inclusions equally from all angles prevents directly comparable, accurate description of all parts of the two species so the generic diagnosis and/or species descriptions may be inaccurate in some respects for one or both species. Additional inclusions are necessary to further clarify apparent similarities and differences between
Neanaperiallus
and
Leptoomus
and between
N. defunctus
and
N. masneri
.