A revision of Physotarsus Townes (Hymenoptera: Ichneumonidae: Ctenopelmatinae), with description of 18 new species Author Zhaurova, Kira Author Wharton, Robert text Zootaxa 2009 2207 1 52 journal article 10.5281/zenodo.189753 96bfbb6b-3a6e-4bbb-8985-e57129b08480 1175-5326 189753 Key to species of Physotarsus Townes, 1966 1 Fore wing weakly ( Fig. 9 ) to strongly ( Fig. 44 ) yellowish or brown to black at base, with ( Figs 9 , 32 ) or without ( Figs 46 , 61 ) infumate areas apically .......................................................................................................................... 2 - Fore wing hyaline at base (as in Figs. 13, 15 ) ............................................................................................................. 8 2(1) Fore wing evenly colored, entirely infumate/fuscous ( Fig. 46 ) or pale yellowish-brown ......................................... 3 - Fore wing not evenly colored (as in Figs 44 , 61 ) ....................................................................................................... 4 3(2) Fore wing pale yellowish brown .......................................................................................... P. albus Zhaurova , n. sp. - Fore wing dark, entirely fuscous .................................................................................. P. m e l i p e n n i s Zhaurova, n. sp. 4(2) Basal part of fore wing dark brown to black, apical part variable ( Figs 44 , 61 ) ........................................................ 5 - Basal part of fore wing yellowish, apex infumate (as in Figs 9 , 32 ) .......................................................................... 6 5(4) Mesosoma dorsally, excluding pronotum, orange; hind femur and tibia orange ..... P. maculipennis ( Cresson, 1874 ) - Mesosoma dorsally, excluding pronotum, dark brown to black; hind femur and tibia black ....................................... ....................................................................................................................................... P. varicornis ( Cameron , 1886 ) 6(4) Mesosoma with yellow and black markings, face quite densely punctate medially ( Fig. 33 ) .................................... ....................................................................................................................................... P. flavipennis Zhaurova, n. sp. - Mesosoma entirely yellowish to light orange, face very lightly evenly punctate ( Fig. 28 ) ........................................ 7 7(6) Frons and first tarsomere of hind leg orange ............................................................................ P. adriani Gauld, 1997 - Frons and first tarsomere of hind leg black ............................................................................ P. castilloi Gauld, 1997 8(1) Epicnemial carina strongly turning towards anterior margin of mesopleuron, reaching it or nearly so .................... 9 - Epicnemial carina largely parallels anterior margin of mesopleuron ....................................................................... 14 9(8) T1 with apical third white, otherwise largely dark brown; ocellar field entirely black .. P. oculatus Zhaurova , n. sp. - T1 either uniformly colored or dark apically, at most with narrow white apical margin; ocellar field variable, entirely black in some species, not in others .......................................................................................................................... 10 10(9) T1 entirely yellowish, orange, or light brown, color even ..................................................................................... 11 - T1 partly to entirely dark brown .............................................................................................................................. 12 11(10) T1 2.3X as long as broad ....................................................................................................... P. eli ethi Gauld, 1997 - T1 1.2X as long as broad ................................................................................................... P. j a m e s i Zhaurova, n. sp. 12(10) Propodeum brown, at least on anterior half ..................................................................... P. niveus Zhaurova , n. sp. - Propodeum entirely orange ....................................................................................................................................... 13 13(12)Clypeus yellowish orange; hind femur dark brown ......................................... P. leucohypopygus Zhaurova , n. sp. - Clypeus white; hind femur yellowish orange ................................................................. P. glabellus Zhaurova , n. sp. 14(8) Ventral clypeal margin with a prominent angulate central lobe ( Fig. 7 ) ................................................................ 15 - Ventral clypeal margin not as above ......................................................................................................................... 16 15(14) Body entirely yellowish orange with no brown or black markings ..................................... P. bonillai Gauld, 1997 - Mesoscutum and face with dark brown markings; anterior half of the propodeum brownish to black ....................... ............................................................................................................................................ P. claviger Zhaurova, n. sp. 16(14) Mesosoma entirely yellow to orange .................................................................................................................... 17 - Color of mesosoma variable, but at least partly black .............................................................................................. 19 17(16) Petiole dark brown ...................................................................................................... P. glabellus Zhaurova , n. sp. - Petiole either pale yellow or dark orange ................................................................................................................. 18 18 (17) Petiole pale yellow; antenna abruptly changing from pale to dark subapically ............. P. luteus Zhaurova , n. sp. - Petiole dark orange; antenna mostly dark throughout, without abrupt color change ......... P. j a m e s i Zhaurova, n. sp. 19(16) Face quite strongly protruding medially ( Figs 49 , 55 ) ......................................................................................... 20 - Face not to very weakly protruding medially ( Fig. 24 ) ............................................................................................ 23 20(19) Mesoscutum largely impunctate posteriorly, very sparsely punctate anteriorly ............ P. gineus Zhaurova , n. sp. - Mesoscutum densely punctate anteriorly and posteriorly ......................................................................................... 21 21(20) Mesoscutum black with anterolateral margin pale yellowish; fore and mid femora partially black ........................ ............................................................................................................................................. P. t o n i c u s Zhaurova, n. sp. - Mesoscutum black with anterolateral margin and most or all of notaular lines pale to dark yellow; fore and mid fem- ora yellowish ............................................................................................................................................................. 22 22(21) Mesopleuron entirely black; face yellowish laterally, black medially ......................... P. foveatus Zhaurova , n. sp. - Mesopleuron black with yellow markings; face entirely yellowish ............................ P. melotarsus Zhaurova , n. sp. 23(19) Mesopleuron and mesoscutum entirely black ........................................................ P. montezuma ( Cameron , 1886 ) - Mesopleuron usually and mesoscutum always with at least some yellow ............................................................... 24 24(23) Tarsomeres 3 and 4 of hind leg bright yellow, much lighter than tarsomeres 1 and 5. Male subgenital plate slightly emarginate ( Fig. 22 ) ......................................................................................... P. concavus Zhaurova , n. sp. - Tarsomeres of hind leg uniformly orange or brown or grading from orange to brown. Male subgenital plate deeply emarginate ( Figs 25 , 31 , 59 ) ..................................................................................................................................... 25 25(24) Tarsomeres of hind leg distinctly infumate, at least dorsally. Antenna mostly dark brown over apical half, often with somewhat indistinct whitish subapical ring ....................................................... P. emarginatus Zhaurova , n. sp. - Taromeres of hind leg orange. Antenna mostly orange to yellow orange over apical half, with apical flagellomeres contrastingly darker..................................................................................................................................................... 26 26(25) Mesoscutum with a narrow anterolateral yellow margin with yellow extending posteriorly along notaular line ( Fig 26 ); male subgenital plate deeply emarginate, cordate ( Fig. 25 ) ..................................... P. cordatus Zhaurova , n. sp. - Mesoscutum with anterolateral margin usually black, never with yellow on notaular line ( Fig 60 ); male subgenital plate deeply emarginate, inner margin not protruding ( Fig. 59 ) ..................................... P. truncatus Zhaurova , n. sp.