Taxonomic notes on Cyperaceae of Nepal: new records of a genus, six species and other noteworthy species
Author
Bhandari, Prabin
https://orcid.org/0000-0002-0199-8656
State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China & University of Chinese Academy of Sciences, Beijing, 100049, China
Author
Chaudhary, Satyam
Central Department of Environmental Sciences, Tribhuvan University, Kirtipur, Nepal
Author
Neupane, Ajay
Mechi Multiple Campus, Tribhuvan University, Bhadrapur, Jhapa, Nepal
Author
Zhou, Shi-Liang
State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China
slzhou@ibcas.ac.cn
Author
Zhang, Shu-Ren
State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China
srzhang@ibcas.ac.cn
text
PhytoKeys
2021
2021-08-09
180
141
156
http://dx.doi.org/10.3897/phytokeys.180.67634
journal article
http://dx.doi.org/10.3897/phytokeys.180.67634
1314-2003-180-141
14A44A7698BB5CE2975CAB8663D02E16
Fimbristylis salbundia (Nees) Kunth, Enum. Pl. 2: 230. 1837.
Trichelostylis salbundia
Nees, Contr. Bot. India 105. 1834.
Type
.
India
,
Silhet,
N. Wallich
3526 [
lectotype
, designated by
Halder and Dey 2016
, pg. 357, 359: K (K000974061 image!)]
.
Description.
Plant rhizomatous, not tufted. Culm up to 130 cm, 5-angled. Leaf reduced to the bladeless sheath, up to 18 cm, tubular. Involucral bracts setaceous to 1 cm long. Inflorescence a compound anthela. Spikelet ovoid, 3.5-4
x
1.5-2 mm, with spirally arranged glumes. Glumes elliptic-ovoid, 1.8-2
x
1 mm, middle part chestnut brown, margin membranous, 3-veined, apex obtuse to acute, not mucronate. Style 1 mm, trigonal, basally inflated, not ciliate. Stigmas 3, as long as style, plumose. Stamens 3, 2 mm long. Achene obovoid, trigonal, 0.5-0.7
x
0.5 mm, sparsely verruculose with transversely oblong epidermal cells in more than 9 vertical rows on each face. (Fig.
2E
).
Distribution.
Nepal, India, China, Myanmar, Bangladesh, Sri Lanka, Philippines, Vietnam, Thailand, Indonesia and New Guinea.
Ecology.
Grows in marshy areas; 760-835 m elev.
Phenology.
Flowering in July-September; fruiting in October-December.
Specimens examined.
Nepal
,
Dang
:
Tulsipur
, near
Damargau
,
Angare
,
835 m
elev.,
17 Dec 2020
,
B. Subedi
20121704
(KATH)
;
Kaski
:
Pokhara Valley
,
Gunde Lake
,
28°11'30.29"N
,
84°2'21.58"E
,
760 m
elev.,
30 Dec 2020
,
P. Bhandari
&
N.L. Bhandari
20123005
(KATH, TUCH)
.
Note.
The protologue of
Fimbristylis salbundia
[≡
Trichelostylis salbundia
] was based on two collections of Wallich, i.e.
Wallich 3499
and
3526
from
'Nepalia'
and
'Silhet'
, respectively (
Wallich 1828
; Nees 1834). All collections representing
3499
were later annotated as
F. quinquangularis
(Vahl) Kunth., except
3499c
at B, which was
F. salbundia
(Nees) Kunth (
Clarke 1907
). The collection
3499c
at B was destroyed in 1943, during the Second World War (
Halder and Dey 2016
). Subsequently, the occurrence of
F. salbundia
was not reported in the published works (
Koyama 1978
;
Press et al. 2000
;
Rajbhandari and Rai 2017
;
Shrestha et al. 2018
;
POWO 2019
;
Govaerts et al. 2021
). The rediscovery of
F. salbundia
after 200 years confirms the occurrence of this taxon in Nepal.
Fimbristylis salbundia
is very similar to
F. quinquangularis
, but can be distinguished, based on the nature of its leaf sheaths and achene character.
Fimbristylis salbundia
is characterised by the presence of bladeless sheaths and sparsely verruculose achene, surface pitted with more than nine vertical rows of transversely oblong epidermal cells. However,
Fimbristylis quinquangularis
has leaf sheaths with blades and densely verruculose achene with up to six vertical rows of transversely linear-oblong epidermal cells.