Taxonomic remarks, phylogeny and evolutionary notes on the leaf beetle species belonging to the Cryptocephalus sericeus complex (Coleoptera: Chrysomelidae: Cryptocephalinae)
Author
Sassi, Davide
text
Zootaxa
2014
3857
3
333
378
journal article
10.11646/zootaxa.3857.3.2
9682bd87-ff66-4cb6-b697-9a53822a5df0
1175-5326
231616
55FBFBCF-F9AF-4EAF-A74A-9A81E6A381B8
Cryptocephalus azurescens
Escalera, 1914
Cryptocephalus azurescens
Escalera, 1914
: 514
.
Type
locality: Telouet,
Morocco
.
Lectotype
(present designation) in MNCN.
Cryptocephalus azurescens
Escalera, 1914
and
C. teleuticus
Escalera, 1914
are poorly known endemic species from the Moroccan Atlas (
Lopatin
et al.
, 2010
).
Escalera (1914)
described them based on specimens collected by himself in Glaui (now Telouet, near Ouarzazate). He separated the two species on the basis of ambiguous and generally unstable characters such as the dorsal colour pattern and the pronotal punctation.
Kocher (1959)
, who studied the
type
series of the two species, kept them apart, but reported that the specimens of
C. telueticus
from Central Atlas examined by him had pronotal punctation denser and strigose, which is precisely the aspect used by Escalera for characterizing
C. azurescens
. He also acknowledged that in the latter species the colour is very variable, emphasizing the presence of several aberrations (
Kocher, 1953
). Besides, Kocher reaffirmed the validity of the separation of the two taxa basing his statement on the analysis of an "aberrant" specimen, which he initially attributed to
C. telueticus
(
Kocher, 1953
)
. Subsequently, however,
Kocher (1959)
assigned this specimen to a chromatic form (“ab.
vaucheri
”) described by
Pic (1950)
and attributed by himself to
C. azurescens
! Thanks
to the courtesy of Dr. Mercedes París (MNCN) I had the opportunity to study the
type
series of the two species, comparing them with several specimens corresponding to the description of Escalera, from Oukamedien, about 60 kilometers West from the
type
locality and from where the presence of
C. azurescens
(
Kocher, 1953
)
has already been reported. I can confirm both the great variability of colour pattern—metallic blue tints and reddish on dorsal surface are distributed in widely differing proportions—and the instability of the punctation of the pronotum. Besides, the shape of the aedeagal median lobe is not significantly different. For these reasons I propose the following synonymy:
C. telueticus
Escalera, 1914
n
. syn. of
C. azurescens
Escalera, 1914
. In order to clarify taxonomic ambiguities I have designated
lectotypes
of both species, choosing specimens that best match Escalera's original description. Aedeagal shape of the
lectotype
of
C. azurescens
is figured (fig. 1).
Material examined:
Type
series of
C. azurescens
. Male: Glaui (handwritten, white label) /
azurescens
(handwritten, white label) / sintipo (printed, red label) /
MNCN
cat. Tipos N° 2234 (partly printed, red label) /
MNCN
_Ent N° Cat. 78236 (printed, grey label) /
Cryptocephalus azurescens
Escalera, 1914
LECTOTYPUS
D. Sassi des. (printed, red label).
Male: Glaui (handwritten, white label) / grupo de
baeticus
nouv (handwritten, white label) /
azurescens
(handwritten, white label) / sintipo (printed, red label) /
MNCN
cat. Tipos N° 2234 (partly printed, red label) /
MNCN
_Ent N° Cat. 78235 (printed, grey label) /
Cryptocephalus azurescens
Escalera, 1914
PARALECTOTYPUS
D. Sassi des. (printed, red label).
Type
series of
C. telueticus
. Male: Glaui (handwritten, white label) / grupo de Pareci [?] bleu (handwritten, white label) / telueti (handwritten, white label) / sintipo (printed, red label) /
MNCN
cat. Tipos N° 2235 (partly printed, red label) /
MNCN
_Ent N° Cat. 78237 (printed, grey label) /
Cryptocephalus telueticus
Escalera, 1914
LECTOTYPUS
D. Sassi des. (printed, red label).
Male: Glaui (handwritten, white label) / sintipo (printed, red label) /
MNCN
cat. Tipos N° 2235 (partly printed, red label) /
MNCN
_Ent N° Cat. 78239 (printed, grey label) /
Cryptocephalus telueticus
Escalera, 1914
PARALECTOTYPUS
D. Sassi des. (printed, red label).
Female: Glaui (handwritten, white label) / telueti (handwritten, white label) / sintipo (printed, red label) /
MNCN
cat. Tipos N° 2235 (partly printed, red label) /
MNCN
_Ent N° Cat. 78238 (printed, grey label) /
Cryptocephalus telueticus
Escalera, 1914
PARALECTOTYPUS
D. Sassi des. (printed, red label).
Besides, 23 further specimens of
C. azurescens
from the environments of Oukaimeden (Haut Atlas,
Morocco
) have been also examined by the author (author's collection).
FIGURES 1–4.
(1) Aedeagus in (a) ventral, (b) dorsal, (c) lateral view:
Cryptocephalus azurescens
lectotype (Glaui [Telouet], Morocco). (2) Aedeagus in lateral view: (a):
C. violaceus scaffaiolus
(Rubiò, Spain); (b):
C. v. violaceus
(Val Gardena, Italy). (3) Aedeagus in ventral view,
C. violaceus scaffaiolus
: (a): (Rubiò, Spain); (b): (Madone d'Utelle, France). (4) Aedeagus in ventral view,
C. v. violaceus
: (a): (Cetinje, Montenegro); (b): (Kovacov, Slovakia). Numbers indicate characters and states. Not drawn to the same scale.
The synonymic summary of
C. azurescens
is as follows:
Cryptocephalus
(
Cryptocephalus
)
azurescens
Escalera, 1914
Cryptocephalus azurescens
Escalera, 1914
: 514
.
Cryptocephalus rugicollis
ab.
vaucheri
Pic, 1950
: 2
.
Cryptocephalus azurescens
ab.
malhommei
Kocher, 1959
: 23
.
Cryptocephalus telueticus
Escalera, 1914
: 515
.
n. syn.
Cryptocephalus telueticus var. pseudobaeticus
Burlini, 1959: 196
.
n. syn.