Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous Author Szadziewski, Ryszard Author Dominiak, Patrycja Author Sontag, Elżbieta Author Krzemiński, Wiesław Author Wang, Bo Author Szwedo, Jacek text Zootaxa 2019 2019-10-24 4688 4 535 548 journal article 25149 10.11646/zootaxa.4688.4.5 c704c5d3-8ef6-48b6-a346-efe5b5e1c032 1175-5326 3517710 F412A472-E501-494B-9BF9-CAF253E9BD92 Culicoides ellenbergeri Szadziewski & Dominiak, sp. nov. Figs 5 A–C urn:lsid:zoobank.org:act: C343228E-064A-4344-A63B-C748F54A468D Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellum with flagellomeres 2–8 spherical, much shorter than the more elongate 9–13, only flagellomere 1 with 7 sensilla coeloconica; distal portion of wing membrane with macrotrichia; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla; proboscis short. Male unknown. Description. Female ( Fig. 5A ). Poorly preserved, distorted, abdomen missing. Eyes bare, separation not visible. Antennal flagellum ( Fig. 5C ) length 0.32 mm ; flagellomere 1 with 7 sensilla coeloconica, flagellomeres 2–13 without sensilla coeloconica, but with distinct sensilla basiconica; flagellomeres 2–8 more or less spherical, much shorter than the more elongate 9–13; antennal ratio 1.13. Palpus ( Fig. 5B ) 5-segmented; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla on ventral surface. Wing ( Fig. 5A ) length 0.61 mm ; costal ratio 0.69; distal portion of wing with macrotrichia in cells r 3 , m 1 and m 2 ; basal radial and cubital cells without macrotrichia. Scutellum with about 8 long marginal setae. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (2) 2.0, TR (3) 1.8. Male. Unknown. Material examined. Holotype female, MAI 5666. Burmese amber, Hukawng Valley , Myanmar ; purchased by Sieghard Ellenberger from Kassel , Germany . Deposited in Museum of Amber Inclusions, University of Gdańsk. Etymology. This new species is dedicated to Sieghard Ellenberger of Kassel, Germany , a generous collector who purchased this biting midge and donated it to the Museum of Amber Inclusions, University of Gdańsk.