Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java
Author
Nurinsiyah, Ayu Savitri
5AFBFEFB-B243-49CC-AACC-248B2F007BC8
Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.
ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id
Author
Neiber, Marco T.
8F6006F5-D880-4841-AC32-B97E18D7BB5C
Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany
mneiber@hotmail.de
Author
Hausdorf, Bernhard
FEDE92C0-108D-4315-8F86-9F2CFF1CB906
Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.
hausdorf@zoologie.uni-hamburg.de
text
European Journal of Taxonomy
2019
2019-05-21
526
1
73
journal article
10.5852/ejt.2019.526
2da65c0a-9c48-43f5-ada3-e088de28dec1
2118-9773
3152217
3FFC527D-E9CA-4D9F-BF59-076FA60171AC
Landouria conoidea
(
Leschke, 1914
)
comb. nov.
Figs 4
,
6
,
53
;
Tables 3–4
Plectotropis conoidea
Leschke, 1914: 212
, figs 8–9 (“Buitenzorg”).
Landouria rotatoria
–
van Benthem Jutting 1950: 461
(in part). (not
Pfeiffer, 1842
)
Diagnosis
Landouria conoidea
is characterized by a small, conical, sharply keeled shell with tubercles all over the shell surface.
Material examined
Syntypes
INDONESIA
•
8 spec.
;
West Java
,
Bogor
;
6°36′ S
,
106°48′ E
;
ZMH
98144
.
Other material
INDONESIA
•
10 spec.
;
West Java
,
Bogor
;
6°36′ S
,
106°48′ E
;
ZMH
98145
.
Description
SHELL (
Figs 4
,
6
;
Tables 3–4
). Conical, with 4.75–5 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side without distinct incised spiral lines, umbilical side with indistinct spiral lines; with tubercles all over shell surface that carry scaly processes or hairs mainly at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled at beginning; aperture almost circular; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus hardly eccentric, wide, comprising 30–43% of shell diameter, hardly obscured by columellar edge.
Figs 6–11.
Shell sculpture of species of
Landouria
from
Java
(umbilical side of shell in 6; apical side in 7–11).
6
.
L. conoidea
(
Leschke, 1914
)
,
West Java
, Bogor (
syntype
, ZMH 98144).
7
.
L. tholiformis
sp. nov.
,
Yogyakarta
, Gunung Kidul, Tepus, near Poktunggal Beach (
holotype
, MZB 17999).
8
.
L. madurensis
sp. nov.
,
East Java
, Madura Island, Sampang, Nepa Monkey lowland rainforest (
paratype
, MZB 19209).
9
.
L. pacitanensis
sp. nov.
,
East Java
, Pacitan, Srau Beach (
paratype
, ZMH 148510).
10
.
L. monticola
van Benthem Jutting, 1950
,
West Java
, Gunung Cikuray (
paratype
, SMF 8880).
11
.
L. smimensis
(
Mousson, 1848
)
,
East Java
, Tosari (ZMA 390881). Scale bars:
1 mm
.
GENITALIA. Unknown.
Remarks
Landouria conoidea
differs from
L. winteriana
, which occurs in the same region in
West Java
, in the smaller shell diameter (
7.6–8.7 mm
vs
9.7–12.6 mm
in
L. winteriana
), the higher shell form (D/H 1.48–1.54 vs
1.55–2.07 in
L. winteriana
), the more pronounced keel and the presence of tubercles all over the shell surface (
Fig. 6
).
With regard to the conical shell form,
L. conoidea
resembles
L. naggsi
sp. nov.
(Fig. 25), from which it differs in the less elevated (D/H 1.48–1.54 vs
1.21–1.37 in
L. naggsi
sp. nov.
), smaller shell (D
7.6– 8.7 mm
vs 11.6–14.0 mm in
L. naggsi
sp. nov.
) with fewer whorls (4.75–5 vs
6.25–7 in
L. naggsi
sp. nov.
), the more pronounced keel and the shell sculpture consisting of tubercles vs longish scales in
L. naggsi
sp. nov.
Distribution
Landouria conoidea
is so far known only from Bogor in
West Java
(Fig. 53).