Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC Landouria petrukensis sp. nov. urn:lsid:zoobank.org:act: 32D7E6E9-561F-4E05-95D2-66295BFD6C5E Figs 19 , 33 , 50 , 52 ; Tables 3–4 Diagnosis Landouria petrukensis sp. nov. is characterized by a rather high, sharply keeled shell with small scales all over the teleoconch, a penis that is divided into a long, broad, cylindrical distal part and a slightly dilated proximal part by a narrow point and a flagellum, which is broadest near its proximal end. Etymology Landouria petrukensis sp. nov. is named after its occurrence in the surroundings of Petruk Cave in Karangbolong. Material examined Holotype INDONESIA • holotype (det. anat.); Karangbolong , way to Petruk Cave ; 7°42′ S , 109°24′ E ; 78 m a.s.l. ; 5 Aug. 2014 ; A.S. Nurinsiyah , F.L.H. Irsyad , F.J. Sari and E. Nurlaela leg.; D = 8.7 mm , H = 5.2 mm ; MZB 19203 . Paratypes INDONESIA – Central Java • 6 spec. ; Karangbolong , near Petruk Cave , agroforestry ; 7°42′17″ S , 109°23′59″ E ; 78 m a.s.l. ; MZB 19273 • 2 spec. ; same data as for preceding; ZMH 133409 • 1 spec. ; same data as for preceding; MZB 19204 • 5 spec. ; Karangbolong , near Petruk Cave , agroforestry ; 7°42′19″ S , 109°24′02″ E ; 78 m a.sl.; MZB 19265 • 4 spec. ; same data as for preceding; ZMH 133417 • 4 spec. ; Karangbolong , wall and surroundings of Petruk Cave ; 7°42′21″ S , 109°24′03″ E ; 104 m a.s.l. ; MZB 19266 • 3 spec. ; same data as for preceding; MZB 19267 • 5 spec. ; Karangbolong , near Petruk Cave , agroforestry ; 7°42′18″ S , 109°24′05″ E ; 138 m a.s.l. ; MZB 19274 • 1 spec. ; same data as for preceding; ZMH 133407 • 23 spec. ; Karangbolong , Adiraja Hill , agroforestry ; 7°42′48″ S , 109°24′05″ E ; 168 m a.s.l. ; MZB 19271 • 4 spec. ; Karangbolong , Adiraja Hill , teak plantation ; 7°42′50″ S , 109°24′05″ E ; 166 m a.s.l. ; MZB 19275 • 3 spec. ; same data as for preceding; ZMH 133414 . Description SHELL (Fig. 50; Tables 3–4 ). Depressed conical, with 4.75–5.5 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles, apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 24–37% of shell diameter, hardly obscured by columellar edge. GENITALIA ( Figs 19 , 33 ; Table 4 ). Atrium short; penis divided by a narrowing into a long, broad cylindrical distal part and a slightly dilated proximal part, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; flagellum moderately long, broadest near its proximal end, where it rapidly tapers, slightly crenated, without node; vas deferens narrow and long; vagina short, distally dilated; oviduct longer, proximally hardly tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina. Remarks With regard to the presence of scales all over the teleoconch, L. petrukensis sp. nov. resembles L. tholiformis sp. nov. (Fig. 51) and L. madurensis sp. nov. (Fig. 57). It differs from both species in the smaller (D 8.1–9.8 mm vs 13.0– 14.4 mm in L. tholiformis sp. nov. and 11.4–12.0 mm in L. madurensis sp. nov. ), more depressed shell (D/H 1.49–1.79 vs 1.77–1.88 in L. tholiformis sp. nov. and 1.75–2.11 in L. madurensis sp. nov. ) with fewer whorls (4.75–5.5 vs 6–6.25 in L. tholiformis sp. nov. and 5.5–5.75 in L. madurensis sp. nov. ), an on average narrower umbilicus (U/D 0.24–0.37 vs 0.32–0.40 in L. tholiformis sp. nov. and 0.38–0.40 in L. madurensis sp. nov. ) and in the penis that is divided by a constriction into a long, broad, cylindrical distal part and a slightly dilated proximal part. We could only investigate the anatomy of the holotype , from Karangbolong in southern Central Java (MZB 19203). There are several samples from other regions of Central Java , West Java and Banten with similar shells. Thus, it is possible that this species is more widely distributed, but the identity of the samples from other regions must be confirmed anatomically. Distribution Landouria petrukensis sp. nov. is known from Karangbolong in southern Central Java (Fig. 52).