Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)
Author
Selnekovič, Dávid
D930D757-D929-4248-A29C-A857E0505345
Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.
david.selnekovic@uniba.sk
Author
Jäch, Manfred A.
D749707A-8823-4110-8D51-BF5AF4E2820F
Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.
manfred.jaech@nhm-wien.ac.at
Author
Kodada, Ján
6E88BFBB-8769-44EC-8285-29E357CEC064
Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.
jan.kodada@uniba.sk
text
European Journal of Taxonomy
2024
2024-09-11
957
1
229
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247
journal article
10.5852/ejt.2024.957.2651
2118-9773
13773385
9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3
Elmomorphus fusiformis
sp. nov.
urn:lsid:zoobank.org:act:
A6F346BC-BEFE-4F95-A988-F025DCDAFF50
Figs 19–20
,
109A
Differential diagnosis
Elmomorphus fusiformis
sp. nov.
(
Fig. 19
) is a remarkably large species (TL
♂♂
3.86–4.13 mm
,
♀♀
3.95–4.29 mm
). It can be characterised by the presence of the dorsal plastron on the entire cranial surface (except for a narrow median area on the vertex), the anterior angles of the pronotum and the lateral sides of the elytra. Large elytral punctures are arranged in nine longitudinal rows on each elytron.
Elmomorphus fusiformis
resembles
E. catenatus
sp. nov.
, which differs in lacking an elytral plastron and in the strial punctures being well delimited and separated in
E catenatus
versus weakly delimited and confluent in
E. fusiformis
.
Fig. 19.
Elmomorphus fusiformis
sp. nov.
, holotype, male (IAECAS), TL: 4.17 mm.
Etymology
The epithet is a Latin adjective in the nominative singular meaning spindle-shaped, referring to the oblong-ovate and strongly convex body shape.
Type material
Holotype
CHINA
–
Yunnan Province
•
♂
; “
CHINA
: NW-Yunnan
15km
N
Lijiang
2800m
,
6.7.1994
leg. JI (16)”;
IAECAS
.
Paratypes
CHINA
–
Yunnan Province
•
9 ♂♂
,
13 ♀♀
; same collection data as for holotype, but some specimens “leg.
H. Schillhammer
”;
CKB
,
NMW
•
1 ♀
; “
China
Yunnan
,
1800m
LIJIANG 23.6.-21.7.
26.53N
100.18E
lgt.
S.Becvar
1992”;
NMW
•
20 ex.
; “
CHINA
:
Yunnan Province
, SHANZHI env.,
22.-24. VI.2007
, JIZU SHAN Mt.,
Zhusheng Si Monastery
,
25°57.7′N
100°23.6′E
,
2180 m
,
J. Hájek
&
J. Růžička
leg. | individually, under stones, on vegetation, in stream; mixed forest (with dominant
Pinus
,
Quercus
, and
Rhododendron
)”;
NMPC
.
Type locality
China
,
Yunnan Province
, Lijiang Autonomous Pref., Lijiang County,
15 km
north of Lijiang City; small valley near the abandoned airport, small stream, 0.5–1.0 m wide, limestone,
2800 m
a.s.l. (
CWBS
60;
Jäch & Ji 1995
).
Description
Measurements (mm): TL:
♂♂
3.86–4.13 (4.00 ±0.11, n =7),
♀♀
3.95–4.29 (4.17 ± 0.14, n=5); PL:
♂♂
0.94–1.04 (0.99 ± 0.04, n= 7),
♀♀
0.96–1.04 (0.99 ±0.03, n=5); PW:
♂♂
1.46–1.67 (1.58± 0.09, n= 7),
♀♀
1.62–1.64 (1.63± 0.01, n =5); EL:
♂♂
2.92–3.09 (3.01± 0.07, n =7),
♀♀
2.99–3.25 (3.17 ± 0.11, n=5); EW:
♂♂
1.82–2.01 (1.93± 0.07, =7),
♀♀
1.92–2.05 (1.99 ± 0.05, n=5); PhL: 0.81– 0.88 (0.85± 0.03, n =7); PrL: 0.37–0.42 (0.39 ±0.02, n =7).
Body oblong-ovate, strongly convex dorsally (
Fig. 19
). Integument black; mouthparts, antennae and tarsi reddish-brown. Pubescence consisting of short decumbent yellowish setae. Dorsal plastron present on cranial surface except for small narrow median area on vertex, on anterior angles of pronotum, and in lateral parts of elytra. Ventral plastron absent on prosternal process, median part of metaventrite, and median part of ventrite 1.
Cranial surface with round setiferous punctures, slightly smaller than an eye facet, separated by 0.5– 1.0 puncture diameter. Labrum transverse, anterior margin slightly emarginate; anterolateral angles rounded; exposed portion microreticulate, with minute round setiferous punctures. Anterior margin of clypeus slightly concave. Eyes oval, interfacetal setae short; ID:
♂♂
0.53–0.60 mm
(0.57 ± 0.03, n= 7),
♀♀
0.56–0.63 mm
(0.61 ± 0.03, n=5); APD/ID:
♂♂
1.70–1.75 (1.72± 0.02, n=7),
♀♀
1.66–1.80 (1.72 ± 0.05, n=5). Antennae 11-segmented. Terminal maxillary palpomere fusiform.
Pronotum transverse, widest at base, PW/PL:
♂♂
1.55–1.64 (1.60± 0.03, n= 7),
♀♀
1.56–1.69 (1.63± 0.01, n=5); surface smooth, large microgranules and coarse setiferous punctures confluent medially; plastron confined to anterior angles. Anterior rim twice as wide as eye facet, interrupted in middle; lateral sides convergent, rounded; anterior angles produced, acute. Prosternal process slightly wider than long; lateral and posterior edges rounded; lateral margins raised, without groups of long setae; median keel moderately raised. Scutellum as long as wide, microreticulate distally, with small round punctures. Metaventral disc flat, without groups of long setae; lateral rims of metaventral process flat. Elytra elongate oval, dorsally convex, widest behind middle, EL/EW:
♂♂
1.53–1.61 (1.56± 0.03, n= 7),
♀♀
1.56–1.63 (1.59 ± 0.03, n=5). Surface weakly microreticulate, small punctures scattered over entire surface, large deep punctures forming nine longitudinal striae, strial punctures separated by about half a puncture diameter. Plastron in two lateral bands, each reaching third stria. Tibiae weakly curved; protibia ca 1.22 × as long as protarsus; PrTL/PL:
♂♂
1.01–1.09 (1.06 ±0.03, n=7),
♀♀
1.03–1.10 (1.06 ± 0.06, n=5). Terminal protarsomere as long as three preceding segments combined.
Fig. 20.
Elmomorphus fusiformis
sp. nov.
, paratype from type locality (CKB), aedeagus.
A
. Lateral aspect.
B
. Ventral aspect. Scale bar: 0.1 mm.
Ventrites covered with plastron, except for median portion of ventrite 1. Ventrite
5 in
males triangularly emarginate at apex; in females rounded, with small glabrous longitudinal keel. Aedeagus (
Fig. 20
): phallobase rather long and slender, basal opening large, PhL/PrL: 2.00–2.30 (2.17± 0.11, n= 7). Parameres moderately curved ventrad, apices rounded in lateral aspect, obliquely truncated in ventral aspect. Penis rounded at apex, sclerotised fibula distinct. Ovipositor: valvifers long, flattened; coxites asymmetrical, right one ca 1.30× as long as left one. Bursa copulatrix without microsclerites.
Secondary sexual dimorphism
Ventrite 5 apically emarginate in males, rounded and with short apical keel in females.
Distribution
China
(
Yunnan
) (
Fig. 109A
).