Description of two new species of Tonnoira Enderlein from caves in the Brazilian Amazon and comments about the taxonomic status of Tonnoira plumaria Quate (Diptera, Psychodidae, Psychodinae) Author Bravo, Freddy Author Chagas, Cinthia Author Alves, Veracilda Ribeiro text Zootaxa 2008 1916 63 68 journal article 10.5281/zenodo.184612 e30cedee-23dc-4989-835c-fee3db1a8d43 1175-5326 184612 Tonnoira robusta Bravo, Alves & Chagas , sp. nov. Figs. 11–20 Type material. BRAZIL , Amazonas, Presidente Figueiredo, Gruta Refúgio do Maragua, 02°03´02.64”S 59°57´47.85”W , 01.IV.2006 , Alves, V. col., holotype male ( INPA ); 1 paratype male, same locality, date, and collector as holotype ( MZUEFS ); 2 paratype males, same locality and collector as holotype , 06.V.2006 ( INPA , MZUEFS ); 2 paratype males, same locality and collector as holotype , 09.II.2006 ( INPA , MZUEFS ); 1 paratype female, same locality and collector as holotype , 09.II.2006 ( INPA ). Description. Male. Head ( Fig. 11 ): eyes separated by 0.5 facet diameters ( Fig. 11 ); interocular suture inverted Y shaped ( Fig.11 ). Antenna: scape cylindrical 1.7X length of pedicel ( Fig. 12 ); pedicel spherical ( Fig. 12 ); flagellum with 14 flagellomeres; all flagellomeres flask-shaped ( Figs. 12, 13 ); first flagellomere same length as second ( Fig. 12 ); apiculus present in flagellomere 14 ( Fig. 13 ); ascoids not visible. Palpus formula = 1.0:2.1:1.9:2.2; last palpomere striated ( Fig. 14 ). Wing ( Fig. 15 ): Sc short; vein R2+3 obsolete, radial fork free; radial fork basal; medial fork complete; R5 ending at wing apex. Male terminalia: hypandrium narrow, diademe-like without ventral expansion ( Fig. 16 ); gonocoxite 0.6X length of the gonostylus ( Fig. 16 ); gonostyli convergent, with base larger than apex ( Fig. 16 ); alveoli present along entire surface ( Fig. 16 ); epandrium rectangular; foramen not observed ( Fig. 17 ); cercopod long, 2.2X length of epandrium, slightly curved ( Fig. 18 ), with 2 tenacula, one apical and the other subapical ( Fig. 19 ); sternite 10 smaller than tergite 10 ( Fig. 17 ); aedeagus asymmetrical, bipartite with left branch smaller than right, curved, pointed at apex ( Fig. 16 ); right branch of aedeagus with rounded apex (fig. 16); with two asymmetrical parameres ending near the apex of the aedeagus apex ( Fig. 16 ); aedeagal apodeme trapezoidal, 0.75x length of aedeagus, wider anteriorly than posteriorly ( Fig. 16 ); gonocoxal apodeme not fused at midline ( Fig. 16 ). FIGURES 11–20. Tonnoira robusta Bravo, Chagas & Alves , sp. nov. Male (11–19), female (20). 11. Head. 12. Antenna: scape, pedicel and basal flagellomeres. 13. Antenna: apical flagellomeres. 14. Palpus. 15. Wing. 16. Male terminalia: dorsal. 17. Male terminalia: tergite and sternite 10 and cercopod. 18. Male terminalia: lateral. 19. Cercopod and apical tenacula. 20. Female terminalia: subgenital plate, genital complex and cerci. d= distance between the base of subgentital plate and apex of apical lobes; S10= sternite 10; T10= tergite 10. Female ( Fig. 20 ). Similar to male except as follows: apical lobes of subgenital plate of females long, rounded, and separated by a shallow apical concavity. Cerci blade-like, 2.2X the total length of genital plate from the anterior border at middle to the lobe apices (distance d). Genital complex small, as illustrated.