Order Rodentia - Family Echimyidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2 1575 1592 book chapter 0-8018-8221-4 10.5281/zenodo.7316535 Proechimys brevicauda (Gunther 1876) [Proechimys] brevicauda (Gunther 1876) , Proc. Zool. Soc. Lond., 1876: 748 . Type Locality: Peru , Dept. de Loreto , Chamicuros (Río Huallago). Vernacular Names: Short-tailed Spiny-rat . Synonyms: Proechimys bolivianus Thomas 1901 ; Proechimys elassopus Osgood 1944 ; Proechimys gularis Thomas 1911 ; Proechimys securus Thomas 1902 . Distribution: Western Amazonia from S Colombia , E Ecuador , E Peru , N Bolivia , and W Brazil . Conservation: IUCN – Lower Risk (lc) as P. brevicauda , P. bolivianus , and P. gularis . Discussion: Listed as a subspecies of longicaudatus by Cabrera (1961:524) and as a member of the longicaudatus group by Patton (1987) , but treated as a species by Patton et al. (2000) . Karyotype has 2n=28-30 and FN=48-50 ( Gardner and Emmons, 1984 ; Patton et al., 2000 ). Likely polytypic, as Gardner and Emmons (1984) suggested gularis is a separate species based on karyotypic grounds and Patton et al. (2000) suggested that elassopus is also distinct, based on morphological and molecular criteria. Anderson (1997) included bolivianus in brevicauda .