New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera) Author Ribeiro, Guilherme C. Author Lukashevich, Elena D. text Zootaxa 2014 3753 4 347 363 journal article 46726 10.11646/zootaxa.3753.4.4 59df87e1-3da4-43ef-8c4c-b8a7cdd580ce 1175-5326 249874 394ADA8F-E7A8-4751-941F-FE8E6A3921A0 Leptotarsus ibericus sp. nov. ( Figs 7, 8 , 22–25 ) Type material. Holotype : LH 28206± (part and counterpart of entire male specimen; preserved structures include the long antenna and terminalia). Paratypes : LH 29995± (part and counterpart of isolated wing) and LH 18122± (part and counterpart of adult). East-central Spain , Las Hoyas; La Huérguina Limestone Formation, Late Barremian, Early Cretaceous. Housed in Museo de Ciencias de Castillia–La Mancha. Etymology. Named after Iberian Mountain Range. Diagnosis. This species differs from species of Leptotarsus from the Cretaceous of Brazil by absence of dark costal field, and from L. buscalioniae sp. nov. by the shorter distal section of M1+2 and distal sections of veins M3 and M4 almost parallel. Description. Head. Head capsule with an anterodorsal protuberance; antennae long and slender, bearing long and slender flagellomeres (total number of flagellomeres unknown); scape stout and long, swollen in middle, 3x longer than pedicel; vestiture of flagellar segments long, delicate and abundant, without visible verticils, length of macrosetae ca. 4.0x the diameter of flagellomere (visible on short portion of antenna); nasus distinct, rostrum ca. 0.5x the total length of head. Thorax stout, as long as wide; transverse suture distinct; praescutal longitudinal stripes clearly delimited. Wing transparent except for dark pterostigma. Sc reaching wing margin at the level of first bifurcation of Rs and beyond level of the tip of Cu; sc-r present, positioned near the tip of Sc; Rs subequal to R2+3 stem; R1 reaching wing margin distal to the level of R2+3 bifurcation; r-r oblique, linking R1 to R2, R2 oblique; medial vein four-branched; discal cell wide; distal sections of veins M3 and M4 subparallel, subequal; long m-cu linking Cu to the base of M4. Abdomen stout, ca. 1.5x longer than thorax; gonocoxite probably conical, as long as wide. Measurements . Antenna length, 19.4. Body length, 7.8. Wing length/width, 6.9–7.9/2.1–2.5.