Revision of Pazala Moore, 1888: The Graphium (Pazala) mandarinus (Oberthür, 1879) Group, with Treatments of Known Taxa and Descriptions of New Species and New Subspecies (Lepidoptera: Papilionidae)
Author
Hu, Shao-Ji
Author
Cotton, Adam M.
Author
Condamine, Fabien L.
Author
Duan, Kuang
Author
Wang, Rong-Jiang
Author
Hsu, Yu-Feng
Author
Zhang, Xin
Author
Cao, Jun
text
Zootaxa
2018
2018-06-28
4441
3
401
446
journal article
29785
10.11646/zootaxa.4441.3.1
80ac1017-65a3-434c-8436-e8701f7761df
1175-5326
1301610
16AB230A-AFCD-484B-ABAD-7CEEDD1532B
Graphium
(
Pazala
)
mandarinus
(
Oberthür, 1879
)
(
Figure 25
)
[
Papilio
]
Glycerion
variété géographique
Mandarinus
Oberthür, 1879
, Étud. d’Ent., 4: 115; TL: “Moupin” [= Baoxing, W. Sichuan, China]. Type locality restricted to Moupin by lectotype designation as stated above. The name
mandarinus
is treated as a noun meaning a Mandarin rather than an adjective, as recommended by Gerardo Lamas (pers. comm.) since there is no clear indication in the original description.
FIGURE 25
. Three known subspecies of
G.
(
P.
)
mandarinus
(Oberthür, 1879)
from Sichuan and Indo-China; upperside in the first row, underside in the second row; scale bar = 10 mm. A:
ssp.
kimurai
Murayama, 1982
, ♂, Doi Inthanon, Chiang Mai, N. Thailand; B:
ssp.
kimurai
Murayama, 1982
, ♂, Mt. Ashaemyinanaukmyin, W. Shan State, Myanmar; C: ditto, ♀; D:
ssp.
fangana
(K. Okano, 1986)
, ♂, Doi Pu Meun, Chiang Mai, N. Thailand; E: ditto, ♀; F:
ssp.
mandarinus
, ♂, Baoxing, W. Sichuan. China; G:
ssp.
mandarinus
, ♀, Daba Shan, W. Sichuan. China.
Diagnostic characters:
Size varies among subspecies, the nominate subspecies is the largest,
ssp.
kimurai
very slightly larger than
ssp.
fangana
which is the smallest, and
ssp.
stilwelli
ssp. nov
.
is intermediate and variable in size, some specimens are quite small and some larger. Both wings whitish and less glossy. Forewing, termen straight or slightly concave in the middle; greyish scales scattered between the 8th and 9th forewing bands. Hindwing, the “8”-shaped twisted discal black bands complete on the underside.
Male genitalia
(
Figure 26
): In total male genitalia of
19 specimens
, comprising two individuals of
ssp.
fangana
, five individuals of
ssp.
kimurai
, six individuals of
ssp.
stilwelli
ssp. nov
.
, and six individuals of
ssp.
mandarinus
, were dissected. Moderately to heavily sclerotised. Ring wavy in the upper half; saccus well developed; distance between the base of socii
0.50–0.64 mm
(mean = 0.56 ± 0.05,
n
= 19), without significant variation between subspecies. Valve round shaped, the dorsal terminal harpe with almost straight edge and acute but not elongate tip (blunt in
sichuanica
and
hoeneanus
nom. nov., stat. rev.
and elongate in
garhwalica
stat. nov.
); the dorsal subterminal harpe moderate in size (similar to that of
daiyuanae
sp. nov.
), isolated or reaching the base of the tip of the dorsal terminal harpe (joining the tip of dorsal terminal harpe in
garhwalica
stat. nov.
; smaller and completely isolated in
paphus
stat. nov.
,
sichuanica
, and
hoeneanus
nom. nov., stat. rev.
; larger in
confucius
sp. nov.
), the medial harpe straight, the dorsal projection finger-shaped (some with blunt or serrate end) (similar to that of
paphus
stat. nov.
and
daiyuanae
sp. nov.
; flat in
garhwalica
stat. nov.
,
sichuanica
, and
hoeneanus
nom. nov., stat. rev.
); with or without a small tooth in the middle of the medial harpe.
Female genitalia
(
Figure 27
): In total female genitalia of
4 specimens
(including two
ssp.
kimurai
, one
ssp.
fangana
, and one
ssp.
mandarinus
) were dissected, the general characters among subspecies are consistent. Lamella postvaginalis small; lamella antevaginalis broad horizontally, lined with weakly sclerotised longitudinal striae; ostial lobe heavily sclerotised, forming a triangular lobe with sharp end in lateral view, while the posterior margin shallowly concave in the middle, with a small short spur at each tip.