Systematics of the Protohermes costalis species-group (Megaloptera: Corydalidae) Author Liu, Xingyue Author Hayashi, Fumio Author Yang, Ding text Zootaxa 2007 1439 1 46 journal article 10.5281/zenodo.175948 02bb7dce-7590-4f45-8aa9-c6280f2e8b36 1175-5326 175948 Protohermes niger Yan g & Ya n g ( Figs. 16 , 95–100 ) Protohermes niger Yang & Yang, 1988 . Zool. Res. 9 (1): 51. Type locality: Yunnan (Fengqing, Lincang). Diagnosis . Head with one pair of blackish markings on lateral margins; pronotum dark, with several indistinct yellowish brown markings; wings grayish brown with several whitish markings. Body length 27–28 mm ; forewing length 35–36 mm , hindwing length 32–33 mm . Head ( Fig. 95 ) yellow or yellowish brown with small post-ocular spine, vertex with one pair of blackish markings on lateral margins. Compound eyes brown; ocelli yellow, darkly margined medially. Median ocellus slightly transverse; posterior pair widely apart, but distance between them slightly shorter than that between antennal bases. Antennae black with scapes and pedicel yellowish brown. Mouthparts yellow; mandible distally black. Prothorax ( Fig. 95 ) dark; pronotum with yellowish brown anterior margin, and with one slender yellow brown spot near each anterolateral corner and one long hook-shaped yellow brown marking following the former spot; central marking cuneiform, yellowish brown, extending to middle. Meso- and metathorax brown, each dorsally with one pair of brownish spots on lateral sides. Thoracic pilosity yellow, much longer on meso- and metathorax. Legs pale brown with dense pale yellowish short setae, tarsal claws reddish brown. Forewings ( Fig. 16 ) grayish brown, with 1 large and 2 small whitish marking at base, 6–7 whitish markings at middle and 1 round whitish marking on apical 1/3; costal area brown. Hindwings grayish brown, with 1 indistinct round whitish marking at base, 1 indistinct round whitish marking at middle and 1 distinct round whitish marking on apical 1/3. Veins dark, except pale yellow on whitish markings. Rs 7 to 8-branched, last branch bifurcate; 6–9 crossveins between R1 and Rs; M1+2 4-branched, M3+4 2-branched; 1A 3-branched. FIGURES 95–100. Protohermes niger Yang & Yang. 95. Male head and pronotum, dorsal view. 96. Male genitalia, dorsal view. 97. Male ninth sternum, ventral view. 98. Male ninth gonostylus, caudal view. 99. Male tenth tergum, ventral view. 100. Male tenth sternum, ventral view. Scale lines = 1.0 mm. Abdomen brown with yellowish short setae. Male ninth tergum ( Fig. 96 ) subquadrate, with trapezoidal anterior margin and shallowly incised posterior margin. Ninth sternum ( Fig. 97 ) broad, posterior margin with shallow trapezoidal incision, forming two broad acute-angled triangular processes; central portion feebly inflated. Ninth gonostylus ( Fig. 98 ) unguiform and strongly curved dorsomedially. Tenth tergum ( Fig. 99 ) short and subcylindrical, ventrally with one well developed tufted tubercle at posteromedial corner; posterolateral corner not produced; tip slightly incised and densely setose. Tenth sternum ( Fig. 100 ) arched with anterior margin feebly elevated; posterior incision triangular and extremely small; lateral lobes thick digitiform and slightly incurved. Female unknown. Material examined . Holotype ɗ, CHINA : Yunnan, Fengqing, Sanchahe, 1200 m , 24.VII.1980 ( CAU ). Paratypes : ɗ, CHINA : same data as holotype ( CAU ); ɗ, CHINA : Yunnan, Lincang, 1750 m , 4.VII.1980 , Qiaosheng Li ( CAU ); ɗ, CHINA : Yunnan, Fengqing, Sanchahe, 1400 m , 20.VII.1980 ( CAU ). ɗ, CHINA : Yunnan, Tengchong, Houqiao, 1700 m , 2.VI.2005 , Hao Huang ( CAU ); ɗ, CHINA : Yunnan, Tengchong, Jietou, 1500 m , 17.V.2006 , Ping Zhao ( CAU ); ɗ, CHINA : Yunnan, Nanjiangonglang, 9.VII.1981 , Dazhi Dong (KIZCAS); 2 ɗ, CHINA : Yunnan, Baoshan, Wayao, 7.VI.1983 , Dazhi Dong (KIZCAS); ɗ, CHINA : Yunnan, Lincang, 20.VI.1979 , Yunzhen Wang (KIZCAS); ɗ, CHINA : Yunnan, Lincang, 4.VII.1981 , Dazhi Dong (KIZCAS); ɗ, CHINA : Yunnan, Lincang, 5.VII.1981 , Pingzhi Yang (KIZCAS); 2 ɗ, CHINA : Lincang, Yongde, 10–23.VII.1981 , Wenguo Yang (KIZCAS). Distribution . China (Yunnan). Remarks . This species is distinguished from the other species of the P. c o s t a l i s group by the distinct blackish pronotum.