Leafy liverworts of Chapada das Mesas National Park: a floristic survey and checklist of the leafy liverworts of Maranhão state, Brazil Author Silva, Jainara Pereira Laboratório de Sistemática Vegetal, Centro de Ciências Agrárias e Ambientais, Universidade Federal do Maranhão, Chapadinha, MA, Brazil Author Oliveira-da-Silva, Fúvio Rubens Programa de Pós-graduação em Ciências Biológicas, Botânica Tropical, Universidade Federal Rural da Amazônia / Museu Paraense Emílio Goeldi, Belém, PA, Brazil • FROS: oliveira. fuvio @ gmail. com https: // orcid. org / 0000 - 0002 - 4871 - 6740 &): 479 - 495. https: // doi. org / 10.15560 / 17.2.479 Author Ilkiu-Borges, Anna Luiza Museu Paraense Emílio Goeldi, Belém, PA, Brazil • ALIB: ilkiu-borges @ museu-goeldi. br https: // orcid. org / 0000 - 0002 - 1266 - 7211 Author Fernandes, Rozijane Santos Laboratório de Sistemática Vegetal, Centro de Ciências Agrárias e Ambientais, Universidade Federal do Maranhão, Chapadinha, MA, Brazil text Check List 2021 2021-03-11 17 1 479 495 journal article 10.15560/17.1.479 1809-127X 5380155 ** Cheilolejeunea acutangula (Nees) Grolle Figure 3E Material examined. BRAZIL • Maranhão , município de Carolina , Parque Nacional Chapada das Mesas , Riacho na RPPN da Mansinha , corticícola em mata de galeria; 07°08′S , 047°26′W ; alt. 286 m ; 3 Feb. 2020 ; JAS Silva 442; CCAA 2330 . Identification. Plants pale green. Ventral merophyte of 2 cells wide. Leaves subimbricate, ovate, apex acute to short acuminate, incurved, margin entire to slightly crenate, dorsal and ventral margin curved; median cells isodiametric, ca. 12‒20 µm, trigones large, dorsal surface of the lobe slightly papillose. Lobules ovate, 1/4 lobe length, apical tooth long, falcate. Underleaves distant, bifid, ovate to orbicular, larger than long or as large as long, 2‒3 × stem width, base cuneate, apex acute, margin ± entire or with a lateral angular tooth. Gynoecia with a pycnolejeuneoid innovation, usually sterile. Perianth obovate, 5-keeled. Cheilolejeunea acutangula is recognized by leaf lobe apex acute to short acuminate, leaf cells with large trigones and dorsal surface of the lobe slightly papillose. This species is commonly reported for xeric environments. Description and illustration. Ilkiu-Borges and Oliveira-da-Silva (2018: 995–997 , fig. 3a–c). * Cheilolejeunea adnata (Kunze) Grolle var. adnata Figure 3F Material examined. BRAZIL • Maranhão , município de Carolina , Parque Nacional Chapada das Mesas , Mata do Buenos Aires , epíxila, na mata ciliar; 06°55′S , 047°22′W ; alt. 174 m ; 4 Feb. 2020 ; JAS Silva 455; CCAA 2350 . * Cheilolejeunea adnata var. autoica Gradst. & Ilk. -Borg. Material examined. BRAZIL • Maranhão , município de Carolina , Parque Nacional Chapada das Mesas , Rio Farinha , corticícola, mata de galeria, Mansinha; 06°59′S , 047°09′W ; alt. 195 m ; 6 Jun. 2018 ; JAS Silva 334; CCAA 1754 .