The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae) Author Alberto Ballerio Author Munetoshi Maruyama text Zookeys 2010 34 77 104 journal article http://dx.doi.org/10.3897/zookeys.34.268 553C91CE-FEA1-4867-929B-2865647E4B2E Besuchetostes howdeni Paulian, 1979 Known distribution: West Malaysia (Pahang) (Paulian 1979). Remarks. This is a flightless species known up to now by the type series only. Examination of the type series revealed that this species does not belong to Besuchetostes Paulian, 1972 (see below, under B. jaccoudi , for more details).