The Type Specimen Of Debeya (Dewalquea) Haldemiana Rediscovered
Author
Halamski, Adam T.
Author
Kvaček, Jiří
National Museum, Department of Palaeontology, Václavské nám. 68,115 79 Praha 1, the Czech Republic; e-mail: jiri. kvacek @ nm. cz
text
Acta Musei Nationalis Pragae Series B
2013
2013-09-30
69
1 - 2
83
86
journal article
10.5281/zenodo.13191046
2533-4069
13191046
Class
Dicotyledoneae
DE
CANDOLLE
1819
Subclass, order, and family incertae sedis
Informal group
Debeya
sensu
KRASSILOV
,
LEWY
,
NEVO
et
SILANTIEVA
2005
1873
1880
Text-figs 1A–G
Dewalquea haldemiana
DEBEY
ex
SAPORTA
et
MARION
(
Araliophyllum haldemianum
DEBEY
in litt.) p. 60; pl. 7, figs 1–2.
Dewalquea haldemiana
DEBEY
ex
SAPORTA
et
MARION
var.
latifolia
HOSIUS
et
VON
DER
MARCK
p. 173; pl 35, fig. 114, pl. 34, fig. 115.
1880
1889
2013
Dewalquea haldemiana
DEBEY
ex
SAPORTA
et
MARION
var.
angustifolia
p. 173; pl 33, figs 116, 117, pl. 34, figs 118-121.
Dewalquea haldemiana
(
DEBEY
ex
SAPORTA
et
MARION
)
ROEMER
, p. 143, pl. 12, figs 2, 3, 4
Debeya haldemiana
(
DEBEY
ex
SAPORTA
et
MARION
)
KNOBLOCH
; Halamski p. 422; figs 7B, C, 8C, 9C, D, F, 10C [
ubi syn.
].
L e c t o t y p e d e s i g n a t e d h e r e: MGUWr 7536p (text-figs 1A–G).
T y p e l o c a l i t y: Haldem, Westphalia,
Germany
.
T y p e h o r i z o n: Campanian, Late Cretaceous.
E m e n d e d d i a g n o s i s. Leaf compound, pedate, petiolate, with 6–7 petiolulate to subsessile linear leaflets; primary venation pinnate, secondary venation brochidodromous; leaflet apices acuminate; tertiary venation ramified.
D e s c r i p t i o n. The
lectotype
shows a compound, pedate, petiolate leaf composed of six leaflets (text-fig. 1
A
). Petiole
28 mm
long and
2.5 mm
wide; inflated proximally,
3.7 mm
wide. Petiolules very short (text-fig.
C
), bases of the leaflets decurrent; sometimes blades begin so closely to the insertion point of the petiolule that the leaflet is subsessile. Leaflets are linear, the median one
210 mm
long, the lateral ones
200 mm
long, up to
13 mm
wide; margins entire, parallel in the median region; apex acuminate. Venation is pinnate consisting of strong primaries. Secondaries are brochidodromous, subopposite, excurrent, originating at an angle of 40–70
o
, first running towards the margin, then subparallel to it, at least looping and joining the higher veins at 1/8 of the blade width (text-figs 1E-G); secondary spacing slightly decreasing or increasing distally depending on the leaflet considered. Intersecondaries are absent. Tertiary veins are poorly preserved, apparently forming a ramified pattern
.