The velvet spiders: an atlas of the Eresidae (Arachnida, Araneae)
Author
Miller, Jeremy A.
Author
Griswold, Charles E.
Author
Scharff, Nikolaj
Author
Řezac, Milan
Author
Szűts, Tamas
Author
Marhabaie, Mohammad
text
ZooKeys
2012
195
1
144
http://dx.doi.org/10.3897/zookeys.195.2342
journal article
http://dx.doi.org/10.3897/zookeys.195.2342
1313-2970-195-1
Dresserus Simon
Dresserus
Simon 1876
: LXXXVII [87]. Type species
Dresserus fuscus
Simon, 1876.
Note.
Dresserus
contains 24 recognized species from eastern and southern Africa (
Platnick 2011
). We examined specimens of multiple unidentified
Dresserus
species. O
bservations
of the male plus the female spinneret spigot morphology is based on specimens from Mazumbai, West Usambara Mts., Tanzania (CASENT 9025746, CAS and CASENT 9025747, CAS); other observations of the female based on specimens from Klein Kariba (CASENT 9025745, CAS) and Drummond (CASENT 9037024, CAS), South Africa; other observations of the male based on specimens from Manga Forest Reserve, Tanzania (Frontier Tanzania, ZMUC), Hluhluwe Game Reserve, South Africa (TM 19739, TMSA), and Witbank, South Africa (TM 19738, TMSA).
Diagnosis.
Distinguished from other eresids except
Gandanameno
by the position of the PLE which are both advanced (<0.28) and widely spaced (PER/AER> 0.95; Fig. 8J); other eresids with advanced PLE (e.g.,
Stegodyphus
, some
Paradonea
) have them closer together (PER/AER <0.90) than the ALE (Figs 10B, 11B, D, F, H, J, L). Generally distinguished from other eresids by the distinctive 4-part cribellum (Fig. 36F), although some
Gandanameno
specimens show signs of this characteristic. Males distinguished from other eresids by the prominent tubercles bearing the ALE (Figs 8J, 33A), which are much more pronounced from those in other genera with ALE tubercles (e.g.,
Stegodyphus
, some
Paradonea
). Further distinguished from other eresids except
Gandanameno
by the more or less ventral-dorsal axis of the palpal bulb with the embolus encircling the ventral part (Figs 12
G-I
, 33
I-K
, 34
A-D
; proximal-ventral in other eresids with the embolus encircling the distal part); distinguished from
Gandanameno
by the smooth conductor (Fig. 34C, D; fringed in
Gandanameno
, Fig. 55C, E). The cephalic region may be wider than long (e.g., TM 19738, TMSA, Frontier Tanzania, ZMUC) or longer than wide (e.g., TM 19739, TMSA). Female further distinguished from other eresids except
Gandanameno
and
Eresus walckenaeri
by the copulatory openings, which are broadly separated by hirsute cuticle (Figs 16C, 37D; separated by a glabrous median lobe in other eresids, e.g., Figs 16B, 29C);
Gandanameno
and
Dresserus
together distinguished from other eresids including
Eresus walckenaeri
by the vulva, which have the spermatheca extending anterior of the spermathecal heads, together subtended by helical copulatory ducts (Figs 16F, 37E; other eresids have the spermathecal head anterior, spermatheca posterior, and copulatory ducts other than helical), and by the subdivided PMS (entire in other eresids) with numerous short, conical, cylindrical(?) gland spigots (Fig. 36C, D; this spigot morphology absent in other eresids); distinguished from
Gandanameno
by the somewhat less prominent paired atria and possibly by having a single loop of the copulatory duct (Fig. 37E; three in
Gandanameno
, although this character has been investigated in few
Dresserus
species).
Distinguishing species.
Thetaxonomic literature on
Dresserus
is fragmentary and rarely comparative. This genus is ripe for revision. We have been unable to confidently assign species names to the specimens used in this study. Lehtinen (1967: 231) indicated that he was working on a taxonomic revision, but none was ever published. According to him, all but one of the described species (including the type species) had been checked and verified as congeneric based on primary types and other material.
Natural history.
Known from Savanna, stony semidesert, and forest habitats. They build a silken tube under stones. Prey is mainly beetles (Milan
Řezac
, personal observa
tion
). Clutches consist of tens of juveniles (Martin Forman, personal observation of
Dresserus kannemeyeri
). Juveniles do not feed on their
mother's
corpse. Mature females can live for several years in captivity and can produce a number of sequential clutches (Martin Forman, personal observation of
Dresserus kannemeyeri
).