Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae) Author Câmara, J. T. Author Rafael, J. A. text Zootaxa 2014 3846 4 502 546 journal article 10.11646/zootaxa.3846.4.2 ea136b1c-dc96-4c41-b078-01ca553453a0 1175-5326 252519 71A80CC3-CDC1-4291-863B-FA7501D60AF5 Opeatocerata curvipenis sp. nov. ( Figs 64–79 ) Diagnosis. Tergites with 3–6 black paramedian subretangular spots ( Figs 66, 67 ); anterior cercus without or with reduced dorsal projection in lateral view ( Figs 68, 69 ); phallus curved ventralward at apex ( Fig. 72 ); genital fork narrow at base and arms longer than base, separated at apex ( Figs 77, 78, 79 ); tergite 10 divided in two subtriangular plates ( Figs 77, 78 ) Description. Male ( Fig. 64 ): Face parallel-sided, dark brown with gray pruinescence visible in ventral view, about 6X longer than lower width. Ocellar tubercle protuberant, dark brown with brown pruinescence; ocelli yellow. Antenna yellow with black bristles; postpedicel about 1.5X longer than pedicel; stylus about 2.5X longer than postpedicel. Proboscis dark yellow, shorter than head height. Thorax dark yellow, shiny ( Fig. 64 ). Legs yellow, except by hind femur and tibia with brown apical ring and all tarsomeres 3–5 black; mid and hind tibia and tarsi with anterodorsal and posterodorsal rows of longer bristles; hind tibia with anteroventral row of longer bristles. Wing ( Fig. 65 ) hyaline, pterostigma brown, conspicuous, about 2.8X longer than wide. Halter yellow. Abdomen yellow; tergites with 3–6 black paramedian subrectangular spots ( Figs 64, 67 ); tergite 8 divided in two subrectangular plates; sternite 8 divided in two trapezoidal plates. Terminalia : Anterior cercus without or with reduced dorsal projection in lateral view ( Figs 66, 68 ); descendant plate subrectangular in posterior view ( Fig. 70 ). Posterior cercus with rounded apex and median subtriangular projection in lateral view ( Figs 66, 68 ); without dorsal projections ( Fig. 69 ). Hypoproct subtriangular in lateral view ( Fig. 68 ) and comma-shaped in posterior view. Epandrial ventral lobe acuminate apically ( Figs 66, 68 ); with spiniform bristles apically and longer bristles mesially ( Fig. 68 ). Hypandrium wider than long, with concave apex ( Fig. 73 ), without long bristles. Phallus wider medially and narrowing towards t apex, with apex curved ventrally, longer than hypandrium ( Fig. 72 ). Subepandrial sclerite wider than long, V-shaped ( Fig. 71 ). Bacilliform sclerite without folds and longer than subepandrial sclerite ( Fig. 71 ). Specimen length : 3.2 mm ; wing length : 3.9 mm . Female ( Fig. 74 ). Similar to male, except frons brown, shiny and wider than face. Face about 4X longer than lower width. Wing ( Fig. 75 ) more infuscated, pterostigma about 3X longer than wide, M1 and M2 veins not reaching wing margin. Terminalia: Tergite 8 subrectangular ( Fig. 78 ). Sternite 8 uniformly wide, straight basally and rounded apically ( Fig. 78 ). Genital fork with narrow base, longer than wide, arms longer than base and separated apically ( Figs 77, 78, 79 ). Tergite 10 undivided, longer than half the length of tergite 8 ( Figs 77, 78 ). Sternite 10 straight at base and bilobed at apex ( Fig. 78 ). Specimen length : 3.3 mm , wing length : 3.9 mm . Geographical distribution . Costa Rica (La Suiza and San Mateo) ( Fig. 211 ). Type material . HOLOTYPE ♂, labelled: La Suiza C[osta] R[ica], Sep[tember]. Pablo Schild / AL Melander Collection, 1961 ( USNM ). Paratypes : Costa Rica , Cartago . Same data ( 1 ♂ , 1 ♀, INPA ). idem , mar ( 1 ♂ , USNM ); idem , jun ( 1 ♂ , INPA ). idem , aug[ust] ( 2 ♂ , 2 ♀, USNM ). Alajuela . San Mateo, Higuito, Pablo Schild Coll. ( 1 ♂ , 1 ♀, INPA ). Pedregosa, D.L. Rounds / AL Melander Collection, 1961 ( 1 ♂ , USNM ). Holotype condition . Abdomen in microtube with glycerin. Etymology . From the Latin curvus (curved) and penis (penis), referring to the ventrally curved apex of the phallus. Variation . Body length ranging from 3.0 to 3.8 mm . Remarks . Opeatocerata curvipenis sp. nov. differs from other species by the following characters: Anterior cercus without or with reduced dorsal projection in lateral view ( Figs 66, 68 ) (in the other species anterior cercus has a well developed dorsal projection), epandrial lobe acuminate ( Figs 66, 68 ) (in other species, the lobe is bilobed, rounded or truncated) and phallus curved downward ( Fig. 72 ) (in other species, the phallus is curved upward).